: Although willows are generally believed to expand population through seed dispersal , there have been some incidents in which a 】 ternative rnetbod for expansion was clearly used . In this study , the authors aimed to under − stand a " seed − ffee " method of willows reproduction , Observations of two s ple populations suggest that wi 且 lows may reproduce and establish new populations in unoccupied sites through branch propagation , a clonal reproduction process by rootilg of deposited branches . As we examined the environmental factors which allowed the new estab − Iishments and the vegetation growth, in those sites where accumu ] ated gravel particleg . were relatively larger had higher皿 umber of individuals ploduced f 士 ol皿 deposited branches instead of growing from seed . 1皿 additio 皿 , larg 巳 branches may have greater chances ofsurviva 且 which may also be an advantage 飴 r healthier growth with this type of reproduction . These results suggest that wiUews may be able to expa 皿 d populations by branches floated f } om up − stream , 働 躍 0 痂 ' 肪 〃OWS ; Pro ρ agation ; Clona〃 e ρ roduct ' 0 η厂Branc 乃, ' Float; Est α biish JOURNAL OF RAINWATERCATCHMENT SYSTEMS / VOL15NO . 12009■ 41
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