No abstract

Phytosauria is a clade of large, carnivorous, semi-aquatic archosauromorphs which reached its peak diversity and an almost global distribution in the Late Triassic (c. 230–201 Mya). Previous phylogenetic analyses of Phytosauria have either focused primarily on the relationships of specific subclades, or were limited in taxonomic scope, and no taxonomically comprehensive dataset is currently available. We here present the most taxonomically comprehensive cladistic dataset of phytosaurs to date, based on extensive first-hand study, identification of novel characters and synthesis of previous matrices. This results in an almost twofold increase in phylogenetic information scored per taxon over previous analyses. Alongside a traditional discrete character matrix, three variant matrices were analysed in which selected characters were coded using continuous and landmarking methods, to more rigorously explore phytosaur relationships. Based on these four data matrices, four tree topologies were recovered. Relationships among non-leptosuchomorph phytosaurs are largely consistent between these four topologies, whereas those of more derived taxa are more variable. Rutiodon carolinensis consistently forms a sister relationship with Angistorhinus. In three topologies Nicrosaurus nests deeply within a group of traditionally non-Mystriosuchini taxa, leading us to redefine Mystriosuchini by excluding Nicrosaurus as an internal specifier. Two distinct patterns of relationships within Mystriosuchini are present in the four topologies, distinguished largely by the variable position of Mystriosuchus. In two topologies Mystriosuchus forms the most basal clade in Mystriosuchini, whilst in the others it occupies a highly derived position within the Machaeroprosopus clade. ‘Redondasaurus’ is consistently recovered as monophyletic; however, it also nests within the Machaeroprosopus clade. The greatest impact on tree topology was associated with the incorporation of continuous data into our matrices, with landmark characters exerting a relatively modest influence. All topologies correlated significantly with stratigraphic range estimates. Topological variability in our results highlights clades in which further investigation may better elucidate phytosaur relationships.

Motion captureDeformed Adding medium Adding high Ground truth markers neutral mesh frequency displacements frequency displacements geometry Figure 1: We synthesize new high-resolution geometry and surface detail from sparse motion capture markers using deformation-driven polynomial displacement maps; our results agree well with high-resolution ground truth geometry of dynamic facial performances. AbstractWe present a novel method for acquisition, modeling, compression, and synthesis of realistic facial deformations using polynomial displacement maps. Our method consists of an analysis phase where the relationship between motion capture markers and detailed facial geometry is inferred, and a synthesis phase where novel detailed animated facial geometry is driven solely by a sparse set of motion capture markers. For analysis, we record the actor wearing facial markers while performing a set of training expression clips. We capture real-time high-resolution facial deformations, including dynamic wrinkle and pore detail, using interleaved structured light 3D scanning and photometric stereo. Next, we compute displacements between a neutral mesh driven by the motion capture markers and the high-resolution captured expressions. These geometric displacements are stored in a polynomial displacement map which is parameterized according to the local deformations of the motion capture dots. For synthesis, we drive the polynomial displacement map with new motion capture data. This allows the recreation of large-scale muscle deformation, medium and fine wrinkles, and dynamic skin pore detail. Applications include the compression of existing performance data and the synthesis of new performances. Our technique is independent of the underlying geometry capture system and can be used to automatically generate high-frequency wrinkle and pore details on top of many existing facial animation systems.

Phytosaurs are a group of carnivorous, semi-aquatic archosaurian reptiles that attained an almost global distribution during the Late Triassic. We here describe a new species of the phytosaur genus Mystriosuchus from the Norian Dachstein Limestone of Austria, from a marine lagoonal depositional environment. The new Austrian material comprises remains of at least four individuals of similar size (c. 4 m in total length) found in association but disarticulated, and includes one complete and two partial skulls and postcrania. All of these specimens apparently represent a single taxon, which is distinguished by numerous anatomical features from the two previously named Mystriosuchus species. Maximum parsimony analysis of a comprehensive morphological dataset provides strong statistical support for the phylogenetic position of the new Austrian taxon in Mystriosuchus, as the sister taxon to a clade comprising M. planirostris and M. westphali. Histological analysis suggests that the Austrian phytosaur specimens represent individuals that were at least eight years old at time of death, but which had not yet reached skeletal maturity. Taphonomic and palaeoenvironmental data suggest that these phytosaurs were living in the marine lagoon in which they were preserved, providing the strongest evidence to date of marine adaptations in phytosaurs.