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Abstract. The Mediterranean Sea is considered a "hot spot" for climate change, being characterized by oligotrophic to ultra-oligotrophic waters and rapidly increasing seasurface temperature and changing carbonate chemistry. Coccolithophores are considered a dominant phytoplankton group in these waters. As marine calcifying organisms they are expected to respond to the ongoing changes in seawater carbonate chemistry. We provide here a description of the springtime coccolithophore distribution in the Mediterranean Sea and relate this to a broad set of in situ-measured environmental variables. Samples were taken during the R/V Meteor (M84/3) oceanographic cruise in April 2011, between 0 and 100 m water depth from 28 stations. Total diatom and silicoflagellate cell concentrations are also presented. Our results highlight the importance of seawater carbonate chemistry, especially [CO 2− 3 ] but also ] in unraveling the distribution of heterococcolithophores, the most abundant coccolithophore life phase. Holo-and heterococcolithophores respond differently to environmental factors. For instance, changes in heterococcolithophore assemblages were best linked to the combination of [CO 2− 3 ], pH, and salinity (ρ = 0.57), although salinity might be not functionally related to coccolithophore assemblage distribution. Holococcolithophores, on the other hand, showed higher abundances and species diversity in oligotrophic areas (best fit, ρ = 0.32 for nutrients), thriving in nutrient-depleted waters. Clustering of heterococcolithophores revealed three groups of species sharing more than 65 % similarities. These clusters could be assigned to the eastern and western basins and deeper layers (below 50 m), respectively. In addition, the species Gephyrocapsa oceanica, G. muellerae, and Emiliania huxleyi morphotype B/C are spatially distributed together and trace the influx of Atlantic waters into the Mediterranean Sea. The results of the present work emphasize the importance of considering holo-and heterococcolithophores separately when analyzing changes in species assemblages and diversity. Our findings suggest that coccolithophores are a main phytoplankton group in the entire Mediterranean Sea and can dominate over siliceous phytoplankton. They have life stages that are expected to respond differently to the variability in seawater carbonate chemistry and nutrient concentrations.

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A Maastrichtian rudist fauna composed of the radiolitids Biradiolites aguilerae Böse, B. Cárdenasensis Böse, Huasteca ojanchalensis (Myers), Tampsia floriformis Myers, and Trechmannites rudissimus (Trechmann), the hippuritids Caribbea muellerriedi (Vermunt) and Praebarrettia sparcilirata (Whitfield) sensu lato, and the plagioptychids Coralliochama gbohemi Böse and Mitrocaprina tschoppi (Palmer) is described from the Cárdenas Formation in San Luis Potosí State, Mexico. Abundant fossil material and excellent preservation of a number of specimens allowed observation of both the internal and external shell characters and their ontogenetic and eco-phenotypic variability. The description of some hitherto insufficiently known species has been enhanced and/or completed, making easier their subsequent identification and allowing their unequivocal generic assignation. Two new genera, Huasteca and Trechmannites, are proposed for two already known species of radiolitids. The Cárdenas Formation exhibits a continuously exposed sequence in the vicinity of Cárdenas. Thus, precise stratigraphic location of all fossil localities and their rudist associations, ranging from the early to the early late Maastrichtian, has been possible.