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Stratigraphic subdivision of the Upper Miocene deposits in the Pannonian Basin has been traditionally based on the endemic mollusc species of Lake Pannon. The cockle species Lymnocardium soproniense Vitális, apparently evolving through a sympatric speciation event in the sublittoral zone of Lake Pannon about 10.2-10.3 Ma, attained wide geographical distribution in the Pannonian basin and thus may serve as a good stratigraphic marker. Lymnocardium soproniense was one of the few large-sized cockles in Lake Pannon, most closely related to its ancestor, L. schedelianum (Fuchs), and to another descendant of the latter, L. variocostatum Vitális. According to the δ 18 O stable isotope record of its shells, the large size of L. soproniense was coupled with an extended life time of more than 10 years, probably reflecting a stable lake environment with increased resource availability and decreased predation. The species lived in quiet offshore conditions, below the storm wave base, where clay was deposited from suspension and the influence of currents was negligible. The base of the Lymnocardium soproniense Zone in the sublittoral deposits of Lake Pannon is defined by the first occurrence of the species, whereas the top of the zone is marked with the base of the overlying Congeria praerhomboidea Zone, defined by the FAD of C. praerhomboidea.

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<p>The integration of sedimentary facies analysis, ichnology and benthic mollusc quantitative paleoecology allowed to interpret factors controlling deposition of the Altavilla Milicia alluvial to marine succession, near Palermo, Sicily. Two major composite units are recognised, separated by an angular unconformity and internally subdivided into elementary depositional sequences. Calcareous nannoplankton biostratigraphy allowed to recognize the upper Pliocene and lower Pleistocene, a time interval that covers the onset of the Quaternary glaciation. The main asset of the succession is driven by tectonic compression and accommodation by transpressional faults, a regime that led to a change in the direction of fluvial sediment delivery, from axial to transverse with respect to the basin elongation. High-frequency eustatic changes have driven the formation of elementary depositional sequences. The upper Piacenzian is characterised by marine bioclastic strata, deposited during maximum flooding intervals of the two large composite units. Mixed carbonate-siliciclastic lithosomes show a good correlation with shallow marine shell-rich detrital carbonates of Northern and Southern Italy and with Mediterranean deep-water sapropel clusters, suggesting common climatic forcing. Some tropical species previously thought to disappear from the Mediterranean at around 3.0 Ma, are instead present in the upper Piacenzian of Sicily. The study suggests that the Pliocene macrobenthos with tropical affinities disappeared from different sectors of the Mediterranean at different times.</p>