A new early dinosaur, Unaysaurus tolentinoi gen. et sp. nov., from the continental Late Triassic red beds of the Caturrita Formation (Carnian-Norian, c. 225 million years old) of southern Brazil is described. U. tolentinoi is represented by a semi-articulated skeleton comprising an almost complete skull, lower jaw and postcranial elements. It differs from all other dinosaurs by several cranial (e.g. developed laterodorsally oriented process formed by frontal and parietal; deep ventral depression on the basisphenoid) and postcranial (presence of a conspicuous blunt ridge running on the lateral surface of the deltopectoral crest of humerus) characters. Unaysaurus represents the first prosauropod grade dinosaur from Brazil and a preliminary phylogenetic analysis indicates it to be closely related to the European Plateosaurus (Plateosauridae). The relationships of the Prosauropoda , however, are still controversial and more comprehensive studies are needed before a clear picture of the evolution and paleobiogeographic distributions of these dinosaurs can be presented.
Present knowledge of Late Triassic tetrapod evolution, including the rise of dinosaurs, relies heavily on the fossil-rich continental deposits of South America, their precise depositional histories and correlations. We report on an extended succession of the Ischigualasto Formation exposed in the Hoyada del Cerro Las Lajas (La Rioja, Argentina), where more than 100 tetrapod fossils were newly collected, augmented by historical finds such as the ornithosuchid
Venaticosuchus rusconii
and the putative ornithischian
Pisanosaurus mertii
. Detailed lithostratigraphy combined with high-precision U–Pb geochronology from three intercalated tuffs are used to construct a robust Bayesian age model for the formation, constraining its deposition between 230.2 ± 1.9 Ma and 221.4 ± 1.2 Ma, and its fossil-bearing interval to 229.20 + 0.11/− 0.15–226.85 + 1.45/− 2.01 Ma. The latter is divided into a lower
Hyperodapedon
and an upper
Teyumbaita
biozones, based on the ranges of the eponymous rhynchosaurs, allowing biostratigraphic correlations to elsewhere in the Ischigualasto-Villa Unión Basin, as well as to the Paraná Basin in Brazil. The temporally calibrated Ischigualasto biostratigraphy suggests the persistence of rhynchosaur-dominated faunas into the earliest Norian. Our ca. 229 Ma age assignment to
Pi. mertii
partially fills the ghost lineage between younger ornithischian records and the oldest known saurischians at ca. 233 Ma.
The upper Triassic deposits of the Selous Basin in south Tanzania have not been prospected for fossil tetrapods since the middle of last century, when Gordon M. Stockley collected two rhynchosaur bone fragments from the so called “Tunduru beds”. Here we present the results of a field trip conducted in July 2015 to the vicinities of Tunduru and Msamara, Ruvuma Region, Tanzania, in search for similar remains. Even if unsuccessful in terms of fossil discoveries, the geological mapping conducted during the trip improved our knowledge of the deposition systems of the southern margin of the Selous Basin during the Triassic, allowing tentative correlations to its central part and to neighbouring basins. Moreover, we reviewed the fossil material previously collected by Gordon M. Stockley, confirming that the remains correspond to a valid species, Supradapedon stockleyi, which was incorporated into a comprehensive phylogeny of rhynchosaurs and found to represent an Hyperodapedontinae with a set of mostly plesiomorphic traits for the group. Data gathered form the revision and phylogenetic placement of Su. stockleyi helps understanding the acquisition of the typical dental traits of Late Triassic rhynchosaurs, corroborating the potential of hyperodapedontines as index fossils of the Carnian-earliest Norian.
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