The degree of sterility resulting in F l hybrids has been used by several workers while considering the relative affinity between the different varieties and sub-species within a species and that between different species . The authors contend that sterility cannot be used as a criterion and present spikelet sterility data in F l hybrids of inter-varietal, inter-subspecific and interspecific crosses in the cultivated rices to support this contention . They conclude that the evolution of the three sub-species (viz ., japonica, javanica and indica) may have taken place independent of each other from the various ecotypes of the putative species Oryza perennis through introgression although the possibility of the indica giving rise to japonica on the one hand and the javanica on the other cannot be altogether ruled out .

The C-A-P genie complex for anthocyanin pigmentation in the apiculus of japonica rice has been recently found to be also applicable to indica rice. The present investigation reveals a considerable difference in the nature and the number of localization genes. Accordingly, the complementary genie complex for anthocyanin pigmentation in the apiculus of both japonica and indica rices is C-A-PaPb or C-A-PaPc or C-A-PbPc. Any two of the localization genes (Pa, Pb, Pc,) together also act as basic genes for other localization genes governing anthocyanin pigmentation in all other parts of the plant.

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