Walking speeds and walking activities (walking time divided by total time) of Trichogramma brassicae were determined at 12, 16, 20 and 25°C. Walking speed was measured during a 5‐min period, and walking activity over a 4‐day period. Both walking speed and walking activity were strongly influenced by temperature. Walking speed increased linearly with the temperature and was twice as high at 20 as at 12°C. At 25°C, walking activity was high during the whole day, at 20 and 16°C it decreased during the afternoon, whereas at 12°C the wasps became most active only at noon or later. At low temperatures, there was a strong individual variation in walking activity. At 25°C, T. brassicae was active most of the time, so the area searched per time unit can only increase at temperatures above 25°C if walking speed increases. At temperatures below 20°C, searching was more restricted by low walking activity than by low walking speed. Even disregarding other effects of temperature, the reduction in walking speed and walking activity at suboptimal temperatures means that T. brassicae can only search half of the area at 20°C, and only one‐seventh at 15°C that it can search at 25°C.
Effects of learning in two microhabitat specialists, Leptopilina boulardi Barbotin et al. and L. fimbriata Kieffer were compared to previous and new results of learning in the microhabitat generalist L. heterotoma Thomson. Females were given one or more oviposition experiences on hosts in different types of substrate. In all species oviposition experience affected the choice for a substrate, although this effect of learning was considerably less in L. fimbriata compared to the other two species. Patch times, known to be highly determined by experience in the generalist L. heterotoma, were much less flexible in the specialists. L. boulardi and L. fimbriata have fixed patch times on their natural substrate and have variable patch times on other substrates only. In all three species one oviposition affected the choice for a substrate. Additional ovipositions showed no different effect. An accumulative effect of the number of ovipositions on patch times was found in L. heterotoma only. Retention of the learning effect was only studied in L. boulardi, and was shown to be similar to that reported for L. heterotoma, i.e. two to three days. Although learning was found in both the generalist and the specialist species studied, it seems to affect their foraging behaviour differently.
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