2010: Assessing the nesting strategies of Late Cretaceous titanosaurs: 3-D clutch geometry from a new megaloolithid eggsite. Lethaia, Vol. 43, Detailed mapping and excavation at the egg locality of Font del Bullidor (Vallcebre Syncline, NE Iberian Peninsula) document various clutches with megaloolithid (Megaloolithus siruguei) eggs and provide significant information on the mode of incubation of Late Cretaceous titanosaurs. Clutches of eggs are preserved in a greyish lagoonal mudstone deposited overlying a sinuous channel infill. Despite the lithological uniformity of the mudstone, clutch architecture can be inferred from the topographical position of the eggs within some clutches. Mapping methods underscore the importance of documenting the spatial relationship of the egg-bearing bed and the mapping plan, as well as an accurate 3-D coordinate acquisition. Both geometry of the clutches and taphonomic observations on particular eggs indicate that eggs were laid in a shallow pit excavated into the substrate. Most of the eggs were incubated under a cover of sediment before hatching or cracking. This corroborates previous inferences on nesting mode.

A new styracosternan ornithopod genus and species is described based on the right dentary of a single specimen from the Mirambell Formation (Early Cretaceous, early Barremian) at the locality of Portell, (Castellón, Spain). Portellsaurus sosbaynati gen. et sp. nov. is diagnosed by two autapomorphic features as well as a unique combination of characters. The autapomorphies include: the absence of a bulge along the ventral margin directly ventral to the base of the coronoid process and the presence of a deep oval cavity on the medial surface of the mandibular adductor fossa below the eleventh-twelfth tooth position. Phylogenetic analyses reveal that the new Iberian form is more closely related to the African taxon Ouranosaurus nigeriensis than to its synchronic Iberian taxa Magnamanus soriaensis and Iguanodon galvensis. In addition, Portellsaurus sosbaynati is less related to other Iberian taxa such as Iguanodon bernissartensis and Proa valdearinnoensis than to the other Early Cretaceous Iberian styracosternans Mantellisaurus atherfieldensis and Morelladon beltrani. A new phylogenetic hypothesis is proposed that resolves Iguanodon (I. bernissartensis, I. galvensis) with the Valanginian Barilium dawsoni into a monophyletic clade (Iguanodontoidea). The recognition of Portellsaurus sosbaynati gen. et sp. nov. as the first styracosternan dinosaur species identified from the Margas de Mirambell Formation (early Barremian–early late Barremian) in the Morella sub-basin (Maestrat Basin, eastern Spain) indicates that the Iberian Peninsula was home to a highly diverse assemblage of medium-to-large bodied styracosternan hadrosauriforms during the Early Cretaceous.

A new spinosaurid genus and species is described based on the right maxilla and five caudal vertebrae of a single specimen from the Arcillas de Morella Formation (Early Cretaceous) at the locality of Cinctorres (Castellón, Spain). Protathlitis cinctorrensis gen. et sp. nov. is diagnosed by one autapomorphic feature as well as by a unique combination of characters. The autapomorphy includes a subcircular depression in the anterior corner of the antorbital fossa in the maxilla. The new Iberian species is recovered as a basal baryonychine. The recognition of Protathlitis cinctorrensis gen. et sp. nov. as the first baryonychine dinosaur species identified from the Arcillas de Morella Formation (late Barremian) from the same time as Vallibonavenatrix cani, the first spinosaurine dinosaur from the same formation in the Morella subbasin (Maestrat Basin, eastern Spain), indicates that the Iberian Peninsula was home to a highly diverse assemblage of medium-to-large bodied spinosaurid dinosaurs. It seems that spinosaurids appeared during the Early Cretaceous in Laurasia, with the two subfamilies occupying the western part of Europe during this period. Later, during the Barremian–Aptian, they migrated to Africa and Asia, where they would diversify. In Europe, baryonychines were dominant, while in Africa, spinosaurines were most abundant.