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The routes of young, inexperienced pigeons released at four sites up to 13 . 5 km from the loft were recorded with GPS-based tracking devices. The routes were found to differ from those of old, experienced pigeons in several aspects : (1) Although being oriented when departing, young birds show more scatter and larger deviations from the home direction, but usually restrict their flights to a semicircle. (2) They apparently ignore prominent landmarks near the loft that are clearly visible. (3) Their tracks are typically more complex, consisting of a number of distinctive phases where the young birds head in different directions, which results in significantly longer routes, often exceeding the direct home distances more than four times. (4) At the same time, young birds seldom venture further away from the release site than the direct distance to home. Their behaviour can be interpreted as exploration to obtain new information on the distribution of navigational factors to be included in their still rudimentary navigational 'map '. At the same time, their flights seem to include elements of safety, like anchoring the flights around the release site and a sense of distance, which help to reduce the chance of getting lost.

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We analysed the tracks of clock-shifted pigeons from six releases to determine how they cope with the conflict between their sun compass and other navigational cues. Time-lag embedding was used to calculate the short-term correlation dimension, a parameter that reflects the complexity of the navigational system, and with it, the number of factors involved. Initially, while pigeons were still at the release site, the short-term correlation dimension was low; it increased as the birds left the site, indicating that the birds were now actively navigating. Clock-shifted pigeons showed more scatter than the control birds, and their short-term correlation dimension became significantly smaller than that of the controls, remaining lower until the experimental birds reached their loft. This difference was small, but consistent, and suggests a different rating and ranking of the navigational cues. Clock-shifted pigeons do not seem to simply ignore the information from their manipulated sun compass altogether, but appear to merely downgrade it in favour of other cues, like their magnetic compass. This is supported by the observation that the final part of the tracks still showed a small deviation in the expected direction, indicating an effect of clock-shifting until the end of the homing flight.