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In the tribe Adeliini, 45 genera are recognised: 24 endemic to Australia, nineto New Zealand, eight to New Caledonia, and three to Chile, and one(Licinoma Pascoe) that occurs in both Australia andChile. The genera of Australia and Chile are keyed and fully described, andall recognised species of Adeliini from Australia (303) and Chile (12),including synonyms and 66 new combinations, are listed in appendices. Onlycryptic features (defensive glands, stridulatory files, aedeagus, ovipositorand female tract) are described for New Zealand and New Caledonian genera, butall characters are included in an overall data matrix for analysis.The Adeliini are considered to be a tribe separate from Laenini. Therelationships of Phobelia Blanchard, Laenini,Goniaderini and Oncotini are briefly discussed. Four generic names aresynonymised: Achora Pascoe, 1869 =Isopteron Hope, 1840; DystalicaPascoe, 1869 = Adelium Kirby, 1818;Macroperas Carter, 1914 =Daedrosis Bates, 1868; andPseudadelium Kaszab, 1982 =Neoadelium Carter, 1908. New specific synonymiesproposed are Daedrosis rufipes Carter, 1934 =Tetragonomenes ruficornis (Champion, 1894);Adelium sinuaticolle Carter, 1914 =A. hackeri Carter, 1908;Dystalica multilineata Carter, 1937 =Seirotrana strigipennis Bates, 1873; andAdelium delicatulumCarter, 1919 =Adelium panagaeicolle Macleay, 1872. Two new names areproposed for new secondary homonyms: Daedrosis carteri,nom. nov. for D. antennalis Carter, 1920 (not Carter,1914), and Isopteron kulzeri, nom. nov. forCestrinus gracilis Kulzer, 1964 (not Carter, 1939).Eleven new genera and seven new species are described from Australia:Apocryphodes based onA. thompsoni, sp. nov; Bellendenumbased on B. gonyxuthum, sp. nov.;Bolusculus based on B. arcanus,sp. nov.; Diaspirus based onD. bellendenus, sp. nov.;Dicyrtodes based on D. arneius,sp. nov.; Diemenoma based onAdelium commodum Pascoe, 1869;Dorrigonum based onLicinoma umbilicata Carter, 1924;Epomidus, based on E. prionodes,sp. nov.; Monteithium based onM. ascetum, sp. nov.; Nolicimabased on Cardiothorax angusticollis Carter, 1906; andYarranum based onSeirotrana crenicollis Pascoe, 1869. Two new genera andone new species are described from Chile: Penadeliumbased on P. araucanum, sp. nov., andValdivium based onAdelium sulcatulum Fairmaire & Germain, 1860.All world genera but one were used for a phylogenetic reconstruction usingparsimony, which in turn is the basis for a biogeographic analysis. It wasfound to be necessary to divide Australia into two areas: the wet tropics ofnorthern Queensland, which have genera forming a monophyletic group with someof those of New Caledonia, and south-eastern Australia, which includes generaforming monophyletic groups with those of Chile. New Zealand genera do notparticipate in convincing monophyletic relationships with those of any othersingle area.

The present paper describes some aspects of the normal behavior o Canthon pilularius (L.) (C. laevis [Drury]) in the field and is the result o direct observations totalling about 5;7 hours, carried out in four field locations in Florida and Georgia in 95;7 and 96. The. approach of this study is neither ethological nor ecological, but taxonomic. That is to say, certain aspects, of the normal behavior of the scarab were. investigated and quanti/qed with a view to using them as taxonomic characters in comparison with other related species and genera. My data on other species of' the genus Canthon are not yet complete enough to present an interspecific analysis of the behavior of this genus, but are adequate for an intergeneric comparison with European representatives of the ball-rolling genera Scarabaeus, Gymnopleurus, and Sisyphus, which have. been studied in detail by German investigators. Such a comparison is now in preparation by the author and will be published subsequently. Previous literature gives the outlines of the li(e history of C. pilularius (Lindquist, 935; Cooper, 938 Ritcher, 945 Miller, 95;4), but there, are no published observations describing the' behavior sequences seen. Brief notes on the biology of about 25 other species of Canthon, sens.u stricto, have been recorded in the. literature (see von Lengerken, 954, Pereira and Mart[nez, 956, and Halffter, 959, for discussions). This. species is commonly known in the United States. literature as Canthon laevis (Drury). However, it has been known for some time that this is not the. correct name. In a recent revision of the. genus (Halffter, 96I) there is a review of the reasons showing why the name pilularius Linnaeus, I758, is most probably correctly attributed to this species. Furthermore, Lane (95o) shows that even the name hudsonias Forster, 77, has precedence over laevis Drury, t773. 'The present investigation was supported in part by Postdoctoral Research Fellowship No. 12,061 of the

Three biogeographic components with very different histories are represented among the tenebrionids of the Australian arid zone: the Indo-Malayan element, which is related mainly to the Indonesian fauna; the Austral Gondwanan element, mainly related to that of southern South America; and one here called Tethyan, related to the fauna of northern hemisphere arid regions. Indo-Malayan groups appear to have arrived comparatively recently, probably by invasion from the north, and are differentiated within the arid zone mainly to the species level. Austral groups are the most diverse and have radiated extensively to generic level from ancestors inhabiting sclerophyll forest, probably by vicariance, within the present arid zone. Tethyan groups are endemic in the arid zone at tribal level and have no forest-inhabiting relatives anywhere. They have often become myrmecophilous in Australia, and because of their distribution patterns in the northern hemisphere, partial occurrence in coastal dunes, and apparently basal phylogenetic positions are surmised to have descended by vicariance from inhabitants of the coastal sand dunes of the Tethys Sea, probably in the Jurassic before there was an arid zone in Australia.