In C3 plants, carbon isotope discrimination (Δ) has been proposed as an indirect selection criterion for grain yield. Reported correlations between Δ and grain yield however, differ highly according to the analyzed organ or tissue, the stage of sampling, and the environment and water regime. In a first experiment carried out in spring wheat during two consecutive seasons in the dry conditions of northwest Mexico (Ciudad Obregon, Sonora), different water treatments were applied, corresponding to the main water regimes available to spring wheat worldwide, and the relationships between Δ values of different organs and grain yield were examined. Under terminal (post‐anthesis) water stress, grain yield was positively associated with Δ in grain at maturity and in leaf at anthesis, confirming results previously obtained under Mediterranean environments. Under early (pre‐anthesis) water stress and residual moisture stress, the association between grain Δ and yield was weaker and highly depended on the quantity of water stored in the soil at sowing. No correlation was found between Δ and grain yield under optimal irrigation. The relationship between Δ and grain yield was also studied during two consecutive seasons in 20 bread wheat cultivars in the Ningxia region (Northern China), characterized by winter drought (pre‐anthesis water stress). Wheat was grown under rainfed conditions in two locations (Guyuan and Pengyang) and under irrigated conditions in another two (Yinchuan and Huinong). In Huinong, the crop was also exposed to salt stress. Highly significant positive associations were found between leaf and grain Δ and grain yields across the environments. The relationship between Δ and yield within environments highly depended on the quantity of water stored in the soil at sowing, the quantity and distribution of rainfall during the growth cycle, the presence of salt in the soil, and the occurrence of irrigation before anthesis. These two experiments confirmed the value of Δ as an indirect selection criterion for yield and a phenotyping tool under post‐anthesis water stress (including limited irrigation).
The relationship between grain yield and carbon isotope discrimination (Δ) was analysed in wheat grown under different water regimes in the Ningxia Province (north‐west of China). When the association was significant, the relationships between grain yield, Δ and other drought tolerance related traits, such as leaf ash content (ma), chlorophyll concentration (Chl), relative water content (RWC), stomatal conductance (gS) and the ratio of internal CO2 leaf concentration to ambient CO2 concentration (Ci/Ca), were also examined. Using correlation analysis, the relationships were determined during two consecutive years in a set of 20 spring wheat cultivars (landraces, improved varieties and advanced lines) under rainfed and irrigated conditions, including saline conditions. The relationship between Δ and yield within environments highly depended on the quantity of water stored in the soil at sowing, the quantity and distribution of rainfall during the growth cycle, and the irrigation before anthesis. Δ predicted grain yield under limited irrigation (post‐anthesis water stress) but not under pre‐anthesis water stress (rainfed conditions), fully irrigated and saline conditions. Under limited irrigation, grain Δ correlated significantly to grain yield leaf ma at heading and maturity. It also significantly positively correlated to Chl, RWC, gS and Ci/Ca assessed at anthesis. A precise characterization of the timing and intensity of the abiotic constraints experienced by the crop is consequently needed before implementing the use of Δ in wheat breeding programmes.
By evaluating the consistency of the Gassmann theory with various inclusion‐based effective medium theories, we investigate the impact of elastic interactions between ellipsoidal pores on the poroelasticity. To rule out any factors that can violate the Gassmann condition, other than elastic interactions, we first construct idealized models that contain only a single set of isolated, identical, and vertically aligned ellipsoidal pores. The numerical simulation suggests that the periodic distribution of ellipsoidal pores generate uniform pore pressure distribution, whereas random distribution of ellipsoidal pores generates heterogeneous pore pressure distributions. Then we analyze the precise conditions under which the underlying Gassmann relationship is valid for various inclusion‐based models. The results reveal the following: (1) Noninteracting effective medium theories are always consistent with the Gassmann prediction, simply because the elastic interactions are ignored. (2) The elastic interactions between randomly distributed pores cause heterogeneous pore pressure that violates the essential requirement of the Gassmann theory. The differential effective medium and self‐consistent approximation theories corresponding to this model thus are inconsistent with the Gassmann prediction. (3) The elastic interactions between periodically distributed pores cause uniform pore pressure; therefore, the Gassmann condition is fully satisfied. The T‐matrix approach explicitly takes into account such elastic interactions and thus is consistent with the Gassmann theory. It is interesting to notice that on top of other well‐known common types of heterogeneities, like pore structure or fluid heterogeneities, the distribution of pores and its associated elastic interactions can be a separate source of heterogeneity, and this makes Gassmann equations not valid anymore.
Conventional seismic analysis in partially saturated rocks normally lays emphasis on estimating pore fluid content and saturation, typically ignoring the effect of mobility, which decides the ability of fluids moving in the porous rocks. Deformation resulting from a seismic wave in heterogeneous partially saturated media can cause pore fluid pressure relaxation at mesoscopic scale, thereby making the fluid mobility inherently associated with poroelastic reflectivity. For two typical gas‐brine reservoir models, with the given rock and fluid properties, the numerical analysis suggests that variations of patchy fluid saturation, fluid compressibility contrast, and acoustic stiffness of rock frame collectively affect the seismic reflection dependence on mobility. In particular, the realistic compressibility contrast of fluid patches in shallow and deep reservoir environments plays an important role in determining the reflection sensitivity to mobility. We also use a time‐lapse seismic data set from a Steam‐Assisted Gravity Drainage producing heavy oil reservoir to demonstrate that mobility change coupled with patchy saturation possibly leads to seismic spectral energy shifting from the baseline to monitor line. Our workflow starts from performing seismic spectral analysis on the targeted reflectivity interface. Then, on the basis of mesoscopic fluid pressure diffusion between patches of steam and heavy oil, poroelastic reflectivity modeling is conducted to understand the shift of the central frequency toward low frequencies after the steam injection. The presented results open the possibility of monitoring mobility change of a partially saturated geological formation from dissipation‐related seismic attributes.
Diffractions caused by, e.g., faults, fractures, and small-scale heterogeneity localized near the surface are often used in ground-penetrating radar (GPR) reflection studies to constrain the subsurface velocity distribution using simple hyperbola fitting. Interference with reflected energy makes the identification of diffractions difficult. We have tailored and applied a diffraction imaging method to improve imaging for surface reflection GPR data. Based on a plane-wave destruction algorithm, the method can separate reflections from diffractions. Thereby, a better identification of diffractions facilitates an improved determination of GPR wave velocities and an optimized migration result. We determined the potential of this approach using synthetic and field data, and, for the field study, we also compare the estimated velocity structure with crosshole GPR results. For the field data example, we find that the velocity structure estimated using the diffraction-based process correlates well with results from crosshole GPR velocity estimation. Such improved velocity estimation may have important implications for using surface reflection GPR to map, e.g., porosity for fully saturated media or soil moisture changes in partially saturated media because these physical properties depend on the dielectric permittivity and thereby also the GPR wave velocity.
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