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High precision oxygen isotope analyses were made of phosphate extracted from 17 samples of nektonic and benthic fish debris sampled across the stratigraphic Cretaceous-Tertiary boundary in northern Morocco. A refinement of the silver phosphate method was used to isolate phosphate from biogenic materials. Measured ~180 values of 18.6-20.5%o are interpreted as reflecting high-resolution thermal variations that affected the ocean water column of the western Tethys. The warm (27°C) water masses that characterized Maastrichtian times underwent rapid cooling and stabilized at an average temperature of 19°C during the Dano-Montian and Thanetian. This period of constant and cool temperature was followed by a relatively rapid but more gradual warming to about 25°C achieved in the Middle Ypresian. Significant small shifts in 6180 values between nektonic and benthic fauna recorded only during the stages of rapid warming or cooling may correspond to averaged thermal differences within the water column that developed in response to global climatic changes. The indicated temperature distribution could have been caused by thermal changes in the atmosphere rather than some signal carried by deep ocean currents. The oxygen isotope data coupled with previous measurements of REE and eNd~TI on the same samples support the suggestion that paleo-Pacific westward currents progressed as far as the northwestern part of the African platform at the end of the Cretaceous period.

Squaliformes comprise the major proportion of modern deep‐water sharks, yet their fossil history and phylogenetic relationships are still poorly understood. New analyses have been undertaken, however, and new living and fossil species have been discovered during the past 10 years. A cladistic analysis involving 29 dental characters has been made and most living and fossil genera are included. On the basis of their dental morphology, the monophyly of the Squaliformes can be supported if the fossil genus Protospinax is excluded. The traditional phylogenetic positions of most living genera, Protosqualus, Cretascymnus and Eoetmopterus, are confirmed despite the fact that the Oxynotidae, Etmopterinae, Palaeomicroides, Proetmopterus and Microetmopterus have some atypical phylogenetic relationships within the Squaliformes. The addition of the palaeontological data in a phylogenetic tree including fossil and living Squaliformes demonstrates some gaps in the fossil record. Nevertheless, and as a consequence of that stratigraphy‐phylogeny inference, two particular events can be pinpointed in the history of the Squaliformes: the first one occurs after the major Cenomanian‐Turonian anoxic event and the second one after the Cretaceous/Tertiary crisis. The first radiation involves the majority of the living Squaliformes (Somniosinae, Centrophorinae, most of the Etmopterinae, Oxynotinae) in deep‐sea waters, the second, the more epipelagic sharks (most of the Dalatiidae), suggesting a secondary adaptation to more shallow environments.

The exact affinities of the fossil teeth attributed to the devilrays (mobulids) are critical for resolving the debated origin of these giant pelagic rays amongst Myliobatiformes and the timing of their evolution toward planktivory. We performed the first detailed comparative description of teeth belonging to most of the living and fossil mobulids. Based on a survey of living devilrays, three dental morphologies are newly identified as cobblestone tooth plates, comb-like teeth, and peg-like teeth. In addition, all extinct mobulid species are reviewed with comments on their dentition, fossil record, and geographical distribution. As a result, three fossil mobulid taxa are newly described from the Late Eocene of south-west Morocco (Argoubia barbei gen. et sp. nov., Oromobula dakhlaensis gen. et sp. nov., and Eoplinthicus underwoodi sp. nov.). This has permitted the first assessment of the phylogenetic positions of extinct and extant species of mobulids, using cladistic analyses and a combined data set of nondental anatomical characters from the literature and the dental characters defined here. Our new results support the monophyly of mobulids including all living and most extinct species and indicate that mobulids are closely related to rhinopterids. They also indicate that there was a recent split within Mobulidae into the three tooth morphology groups that we describe in this paper. This work provides clues to the evolutionary history of this clade since the Early Eocene, including the gradual lack in tooth interlocking toward the filter-feeding strategy, whereas the preservation of cusped teeth without feeding function in modern filter-feeder mobulids is interpreted as a tool for precopulatory purposes.