The Karoo igneous rocks represent one of the largest continental flood basalt events (by volume) on Earth, and are not normally associated with fossils remains. However, these Pliensbachian-Toarcian lava flows contain sandstone interbeds that are particularly common in the lower part of the volcanic succession and are occasionally fossiliferous. On a sandstone interbed in the northern main Karoo Basin, we discovered twenty-five tridactyl and tetradactyl vertebrate tracks comprising five trackways. The tracks are preserved among desiccation cracks and low-amplitude, asymmetrical ripple marks, implying deposition in low energy, shallow, ephemeral water currents. Based on footprint lengths of 2-14 cm and trackway patterns, the trackmakers were both bipedal and quadrupedal animals of assorted sizes with walking and running gaits. We describe the larger tridactyl tracks as "grallatorid" and attribute them to bipedal theropod dinosaurs, like Coelophysis, a genus common in the Early Jurassic of southern Africa. The smallest tracks are tentatively interpreted as Brasilichniumlike tracks, which are linked to synapsid trackmakers, a common attribution of similar tracks from the Lower to Middle Jurassic record of southern and southwestern Gondwana. The trackway of an intermediate-sized quadruped reveals strong similarities in morphometric parameters to a post-Karoo Zimbabwean trackway from Chewore. These trackways are classified here as a new ichnogenus attributable to small ornithischian dinosaurs as yet without a body fossil record in southern Africa. These tracks not only suggest that dinosaurs and therapsids survived the onset of the Drakensberg volcanism, but also that theropods, ornithischians and synapsids were among the last vertebrates that inhabited the main Karoo Basin some 183 Ma ago. Although these vertebrates survived the first Karoo volcanic eruptions, their rapidly dwindling habitat was turned into a land of fire as it was covered by the outpouring lavas during one of the most dramatic geological episodes in southern Africa.

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Interdune deposits are sensitive to climatic fluctuations and ancient interdune lacustrine systems can reveal the drivers of erg dynamics through time and space. This southern African study details the dynamics of interdune deposition in a vast ancient desert system that was active over Pangaea in the Middle Mesozoic and formed part of one of the largest known sand seas in Earth's history. Focusing on the Sinemurian to Pliensbachian Clarens Formation in the main Karoo Basin (south-west Gondwana), three facies associations were identified that formed in meso-scale and macro-scale lakes amongst aeolian dunes and sand sheets of the Clarens erg system. The northern facies association is restricted to interdune hollows within an actively climbing dune system. High-sediment availability and high-wind speeds limited the space available for these meso-scale lakes to expand. Here, migrating dunes outpaced the water table rise promoting the formation of discontinuous lenses of lake deposits, as is typical in erg centres. The southern facies associations developed both in spatially isolated meso-scale lakes, where flooding was controlled by groundwater inundation (water table rise), as well as in macro-scale lakes, where flooding was related to groundwater inundation and floods (surface runoff) that induced terminal splays. Relative to the north, the southern facies associations, with their more significant thickness and lateral extent, show that, here, the basin was characterized by a higher rate of relative subsidence. Moreover, the southern lake systems were established along the erg's margin, where the interdunes expanded during pluvial phases. Ultimately, the facies associations in the Clarens Formation show that the widespread arid conditions in the Early Jurassic of south-west Gondwana were subjected to prolonged and repeated pluvial phases in an otherwise harsh environment. Given the prominence of Jurassic aeolianites across Pangea, this study provides a southern African perspective on global palaeoerg dynamics and Early Jurassic climatic shifts.

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