The biosphere is changing rapidly (Ellis et al., 2010) through anthropogenic activities that result in fragmentation and loss of natural habitats Harrison et al., 2017). These changes cause increasing threats to biodiversity, especially among species with ecological interactions such as pollinators, influencing the delivery of ecosystem services necessary for the maintenance of both natural and agricultural ecosystems (Brittain et al., 2013;CBD, 2018; IPBES, 2016). Anthropogenic factors such as habitat fragmentation affect the extent of natural habitat (Goulson et al., 2008) and habitat suitability for many species (Tscharntke et al., 2002). Consequently, to preserve local biodiversity and key ecosystem functions, protected areas (PAs) are established (Andam et al., 2008;Geldmann et al., 2013;Mukul et al., 2017). This is essential for the protection of biodiversity in highly diverse yet disturbance-sensitive ecosystems such as the tropical savanna and rain forest vegetation zones of Nigeria, where over half of the primary forest has been lost (FAO, 2015;Mfon et al., 2014). There is evidence of increasing threats to biodiversity from various anthropogenic factors in this region (BESNet, 2019;Kehinde et al., 2014); however, conservation planning has been hindered by a lack of adequate information on
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