No abstract

The Carabus subgenus Hygrocarabus contains two taxa: C. variolosus and C. nodulosus, the species or subspecies status of which is handled far from uniform in the literature. Both taxa show a similar morphology, the shape of the tip of the aedeagus provides a reliable morphological marker for identification. We analysed two mitochondrial gene parts (COI-5’ and COI-3’) and a nuclear one (ITS2). High diversity was found showing specific geographical patterns. Introgressive hybridisation was detected but interpreted not as an argument for subspecies status because high genetic distances indicated that it must have taken place in former times. In a laboratory hybridisation experiment, the male did not accept the female of the other taxon, supporting the conclusion that these are separate species. A series of refuges was expected for the period of ice ages. Although only the taxon C. variolosus is listed in Annex II and IV of the EU Habitats Directive, C. nodulosus also falls under this listing, as at the time of including the species into the Annexes in 2004, the two taxa were considered subspecies and hence the listing would include both, independent of later taxonomic revisions.

In this paper, five new subspecies of Carabus Linnaeus, 1758 (Coleoptera Carabidae) are described and figured. Three are from the southeastern part of Sichuan province (China): Carabus (Apotomopterus) longeantennatus transjinshaensis n. ssp.; Carabus (Apotomopterus) benardi briseis n. ssp.; Carabus (Acoptopterus) vigilax rautenstrauchi n. ssp. The other two are from the Russian Caucasus (Ingushetia and Daghestan): Carabus (Tribax) kasakorum daghestanicola n. ssp.; Carabus (Tribax) kasakorum scintillans n. ssp. Furthermore, Carabus (Apotomopterus) ebian Cavazzuti et Rapuzzi, 2010 is considered a valid species after examination of a male specimen. The male aedeagus is described and figured for the first time.

In this paper the authors record for the first time the invasive species Callidiellum rufipenne(Motschulsky, 1860) (Coleoptera Cerambycidae) for the fauna of the republics of Bosnia andHerzegovina and Slovenia.

Four new subspecies of genus Carabus linnaeus, 1758 (Coleoptera Carabidae) belonging to subgenus Apotomopterus and one new Carabus subspecies belonging to subgenus Pseudocoptolabus are described and figured. Two of these are originally from the southeastern part of Sichuan province (China): Carabus (Apotomopterus) kassandra planielongatus n. ssp. and Carabus (Apotomopterus) benardi alterego n. ssp.; one from Hunan province (China): Carabus (Apotomopterus) cyanipennis atripennis n. ssp.; one from northeast Yunnan province (China): Carabus (Pseudocoptolabrus) taliensis xerophilous n. ssp.; one from Ha Giang province (Vietnam): Carabus (Apotomopterus) tonkinensis hagiangensis n. ssp. Carabus (Pseudocoptolabrus) chortenensis Cavazzuti, 2005 is considered as a valid species after examination of a male specimen from its type locality; the male imago and aedeagus are described and figured for the first time. Carabus (Pseudocoptolabrus) chortenensis mosso deuve et mourzine, 2005 comb. nov. RESULTS Systematics Carabus (Apotomopterus) kassandra planielongatus n. ssp.-Fig. 1-http://zoobank.org/ 67ab6c74-0790-4358-b34a-04558dfd1686 ExaminEd maTERial. Holotype female, China,