The vegetation, environment, and ecological gradients present on three calcareous mires in the South Park valley, Park County, Colorado, were investigated. Vegetation was classified into four habitat classes, nine subclasses, and twelve species associations using two-way species indicator analysis (TWINSPAN). Detrended correspondence analysis (DCA) was used to ordinate vegetation samples along two axes representing the three predominant ecological gradients: water table height, miremargin to expanse, and region. Canonical correspondence analysis (CCA) was used to directly relate local environmental conditions to vegetation. Water table depth, microtopographical development, soil and water pH and nutrient level, soil organic matter, and hydraulic head were significantly correlated with vegetation gradients. The mire soils consist of intermixed areas of organic and mineral soils. Mire soils and water are highly alkaline and nutrient-rich. Mean pore water calcium concentration on these mires is 115 mg/L, electrical conductivity averages 575 µS, and mean pH is 7.4. Owing to these conditions, the fen floras include a number of highly minerophilic species. Based on water chemistry and species criteria, each site was classified as rich to extremely rich fen, with the two fen types mixing in complex patterns according to local environmental conditions. The species Trichophorum pumilum, Salix candida, Salix myrtillifolia, Carex microglochin, Carex viridula, Carex scirpoidea, Eriophorum gracile, Triglochin maritimum, Triglochin palustris, Kobresia myosuroides, Kobresia simpliciuscula, Thalictrum alpinum, Scorpidium scorpioides, Scorpidium turgescens, and Calliergon trifarium were determined to be indicative of extremely rich fen conditions in the southern Rocky Mountains.Key words: Colorado, canonical correspondence analysis, detrended correspondence analysis, extremely rich fen, gradient analysis, mire.

An AMS radiocarbon-dated pollen record from a peat deposit on Mitkof Island, southeastern Alaska provides a vegetation history spanning ∼12,900 cal yr BP to the present. Late Wisconsin glaciers covered the entire island; deglaciation occurred > 15,400 cal yr BP. The earliest known vegetation to develop on the island (∼12,900 cal yr BP) was pine woodland (Pinus contorta) with alder (Alnus), sedges (Cyperaceae) and ferns (Polypodiaceae type). By ∼12,240 cal yr BP, Sitka spruce (Picea sitchensis) began to colonize the island while pine woodland declined. By ∼11,200 cal yr BP, mountain hemlock (Tsuga mertensiana) began to spread across the island. Sitka spruce-mountain hemlock forests dominated the lowland landscapes of the island until ∼10,180 cal yr BP, when western hemlock (Tsuga heterophylla) began to colonize, and soon became the dominant tree species. Rising percentages of pine, sedge, and sphagnum after ∼7100 cal yr BP may reflect an expansion of peat bog habitats as regional climate began to shift to cooler, wetter conditions. A decline in alders at that time suggests that coastal forests had spread into the island's uplands, replacing large areas of alder thickets. Cedars (Chamaecyparis nootkatensis, Thuja plicata) appeared on Mitkof Island during the late Holocene.

The vegetation of a subalpine fen in Colorado was studied. Insight was sought into the community structure and factors influencing species distribution of a vegetation type heretofore undescribed in the southern Rocky Mountains. A vegetational gradient was evaluated using detrended correspondence analysis (DCA). Four types of vegetation were subjectively defined; these same types were distinguished by the DCA. DCA further revealed marked differences in the vegetation occurring on peat hummocks versus in hollows. Species composition was related to environment using canonical correspondence analysis (CCA). Water-table depth, hummock height, shading, groundwater temperature, and conductivity were significantly correlated with species distribution, accounting for 51% of the total species variance. Univariate regression was used to examine how tree density varied with environment. The above factors, except for shading and conductivity, were also significantly correlated with tree density. It is suggested that the peat hummocks that form on this moderate fen provide an environment similar to that of an ombrotrophic bog and that these "miniature bogs" form in areas unable to support expansive bogs. Further, these hummocks provide small-scale environmental heterogeneity that exerts a strong control over species composition that would not be evident in studies based on samples of a large areal extent. Keywords: Colorado, gradient analysis, ordination, heterogeneity, peatlands, phytosociology.

Abstract. The tree population size structure and relationship between tree diameter and age were examined in a subalpine fen and surrounding Picea‐Abies forest in northern Colorado. The fen grades from a sedge fen, through an ecotone, to a treed fen. Tree growth rate varies across the vegetational gradient, with the sedge fen having the slowest growth, and the upland forest having the fastest growth. Differences in growth rate are related to the average size of peat hummocks, with areas containing tall hummocks exhibiting the highest tree growth rates. Size structures display the characteristic reverse‐J distribution generally indicative of stable populations, but forest vegetation is expanding into the open regions of the fen, and within the treed fen an increase in Abies lasiocarpa is occurring. These changes are primarily attributed to a positive feedback situation wherein the fen's surface is built up by peat accumulation. Distinct hummocks form first on the open fen but then coalesce to form raised peat islands in the treed fen. This new substrate provides habitat with a comparatively low water table and allows the growth of mesophytic forest vegetation. A pathway for this vegetational development is proposed.