This long-term experiment in Maine, U.S.A., was designed to provide information on the best silvicultural practices for managing stands of mixed northern conifers in northeastern U.S.A. We evaluated growth and yield and changes in species composition, quality, and structure during the first 40 years of the experiment. Replicated treatments include the selection system, uniform shelterwood, unregulated harvesting, and diameter-limit cutting. The new cohort established under three-stage shelterwood was subsequently left untreated or precommercially thinned. Between-treatment differences in net volume growth were not significant (α = 0.10), though gross volume growth differed significantly for managed vs. unmanaged, selection vs. shelterwood, and shelterwood vs. diameter-limit treatments. A three-stage shelterwood method with precommercial thinning 10 years following final overstory removal resulted in good control of hardwoods and hemlock and a dramatic increase in spruce and fir. The selection system on a 5-year cutting cycle resulted in increased hemlock, spruce, and fir, with a decrease in hardwood species. If the primary goal were production, even-aged management would most likely be preferred. We recommend a two-stage shelterwood method as applied in this experiment with some modification to improve species composition and stand quality. Stand quality (proportion of stand volume in cull trees) and species composition was influenced by treatment.

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Several hypotheses about the relationships among individual tree growth, tree leaf area, and relative tree size or position were tested with red spruce (Picea rubens Sarg.) growing in uneven-aged, mixed-species forests of south-central Maine, U.S.A. Based on data from 65 sample trees, predictive models were developed to (i)estimate the amount of foliage held by individual trees from sapwood cross-sectional area and (ii)define the relationship between stem volume growth and three variables: total foliage area, relative position in the stand, and the degree of past suppression. A model that included variables representing tree size (or relative social position) and degree of past suppression (live branch whorls per unit crown length) indicated that stem volume growth first increased but later decreased over leaf area when other variables were held constant. Growth efficiency declined with increasing tree leaf area, although greater height and diameter enhanced growth efficiency and greater past suppression diminished growth efficiency. The decline in growth efficiency with greater leaf area likely is attributable to one or several of the factors previously identified as contributing to growth declines in mature, even-aged stands.

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