Savanna ecosystems are characterized by the co‐occurrence of trees and grasses. In this paper, we argue that the balance between trees and grasses is, to a large extent, determined by the indirect interactive effects of herbivory and fire. These effects are based on the positive feedback between fuel load (grass biomass) and fire intensity. An increase in the level of grazing leads to reduced fuel load, which makes fire less intense and, thus, less damaging to trees and, consequently, results in an increase in woody vegetation. The system then switches from a state with trees and grasses to a state with solely trees. Similarly, browsers may enhance the effect of fire on trees because they reduce woody biomass, thus indirectly stimulating grass growth. This consequent increase in fuel load results in more intense fire and increased decline of biomass. The system then switches from a state with solely trees to a state with trees and grasses. We maintain that the interaction between fire and herbivory provides a mechanistic explanation for observed discontinuous changes in woody and grass biomass. This is an alternative for the soil degradation mechanism, in which there is a positive feedback between the amount of grass biomass and the amount of water that infiltrates into the soil. The soil degradation mechanism predicts no discontinuous changes, such as bush encroachment, on sandy soils. Such changes, however, are frequently observed. Therefore, the interactive effects of fire and herbivory provide a more plausible explanation for the occurrence of discontinuous changes in savanna ecosystems.

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Hyperspectral remote sensing serves as an effective tool for estimating foliar nitrogen using a variety of techniques. Vegetation indices (VIs) are a simple means of retrieving foliar nitrogen. Despite their popularity, few studies have been conducted to examine the utility of VIs for mapping canopy foliar nitrogen in a mixed forest context. In this study, we assessed the performance of 32 vegetation indices derived from HySpex airborne hyperspectral images for estimating canopy mass-based foliar nitrogen concentration (%N) in the Bavarian Forest National Park. The partial least squares regression (PLSR) was performed for comparison. These vegetation indices were classified into three categories that are mostly correlated to nitrogen, chlorophyll, and structural properties such as leaf area index (LAI). %N was destructively measured in 26 broadleaf, needle leaf, and mixed stand plots to represent the different species and canopy structure. The canopy foliar %N is defined as the plot-level mean foliar %N of all species weighted by species canopy foliar mass fraction. Our results showed that the variance of canopy foliar %N is mainly explained by functional type and species composition. The normalized difference nitrogen index (NDNI) produced the most accurate estimation of %N (R 2 CV = 0.79, RMSE CV = 0.26). A comparable estimation of %N was obtained by the chlorophyll index Boochs2 (R 2 CV = 0.76, RMSE CV = 0.27). In addition, the mean NIR reflectance (800-850 nm), representing canopy structural properties, also achieved a good accuracy in %N estimation (R 2 CV = 0.73, RMSE CV = 0.30). The PLSR model provided a less accurate estimation of %N (R 2 CV = 0.69, RMSE CV = 0.32). We argue that the good performance of all three categories of vegetation indices in %N estimation can be attributed to the synergy among plant traits (i.e., canopy structure, leaf chemical and optical properties) while these traits may converge across plant species for evolutionary reasons. Our findings demonstrated the feasibility of using hyperspectral vegetation indices to estimate %N in a mixed temperate forest which may relate to the effect of the physical basis of nitrogen absorption features on canopy reflectance, or the biological links between nitrogen, chlorophyll, and canopy structure.

The continuity equation, Manning's equation and an empirically determined relationship between channel roughness and the biomass of macrophytes were used to simulate the effects of weed growth in contrasting channels. Two indices of wetted available habitat, velocity and depth were chosen to illustrate the role of macrophyte in maintaining and modifying instream habitat with particular reference to chalk streams. Plant growth maintained depth within the channels and its effect was modified by channel shape and slope. Weed cutting resulted in very sudden changes in depth and velocity and the loss of a large volume of water from the river. The results indicate that macrophyte growth could be used to maintain wetted habitat while allowing more abstraction, but more data are required on the long-term effects of implementing such policies.

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