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The sedimentological and micropaleontological analysis of three mechanical cores in the marshland of Almenara (Valencian Community, Spain) has allowed the reconstruction of the Holocene evolution of this wetland. The cold and dry 8.2-ka event might be represented in Almenara by a massive carbonate precipitation bed, upon which mid- and late-Holocene sediments were subsequently deposited. The direct influence of sea-level changes has been recorded in the two cores (S-4 and S-5) located near the marsh barrier, at 400–450 m from the current coastline. The maximum flood area during MIS 1 (last 11,600 years) is represented in these cores by sediments indicative of different littoral subenvironments (shoreface, foreshore, backshore). These sediments contain typically littoral marine foraminiferal species such as Ammonia beccarii, Rosalina globularis, Asterigerinata mamilla, Adelosina longirostra, Cibicidoides lobatulus, Elphidium macellum, and Bolivina pseudoplicata. The base of these littoral sedimentary materials has been dated as 5480 and 5580 cal. yr BP. At this moment, the inner area (core S-7) was occupied by a restricted oligohaline marsh subject to water-table fluctuations and with scarce individuals of brackish water foraminifera, such as Ammonia tepida, Haynesina germanica, or Cribroelphidium excavatum, that in more recent times (since at least 1700 cal. yr BP) gradually evolved to a palustrine area.