No abstract

The Triassic chronostratigraphic scale was built on two centuries of research on ammonoid biostratigraphy and biochronology. Two Triassic stage bases and all of the Triassic substages are currently defined by ammonoid bioevents. The study of Triassic ammonoids began during the late 1700s, and in 1895, Edmund von Mojsisovics, Wilhelm Waagen and Carl Diener published an essentially complete Triassic chronostratigraphic scale based on ammonoid biostratigraphy. This scale introduced many of the Triassic stage and substage names still used today, and all terminology of stages and substages subsequently introduced has been based on ammonoid biostratigraphy. Early Triassic ammonoids show a trend from cosmopolitanism (Induan) to latitudinal differentiation (Olenekian), and the four Lower Triassic substage (Griesbachian, Dinerian, Smithian and Spathian) boundaries are globally correlated by widespread ammonoid biotic events. Middle Triassic ammonoids have provinciality similar to that of the Olenekian and provide a basis for recognizing six Middle Triassic substages. Late Triassic ammonoids provide a basis for recognizing three stages divided into five substages. The main uncertainty for the future of Triassic ammonoid biostratigraphy is not the decline of the ammonoids as a tool for dating and correlation of Triassic strata but, rather, the dramatic decrease in the number of specialists, due to the lack of replacement of experienced palaeontologists who started their activity in the 1950s and 1960s.

The first abundant, well-preserved fossils of the unusual archosauromorph reptile Trilophosaurus jacobsi Murry are from an Upper Triassic bonebed in the lowermost Trujillo Formation of the Chinle Group in Borden County, Texas. A nearly complete left side of the skull and incomplete but articulated mandible of a juvenile individual demonstrate that Trilophosaurus jacobsi Murry is referable to Trilophosaurus, so the putative procolophonid genus Chinleogomphius is a junior objective synonym of Trilophosaurus. Features of T. jacobsi that diagnose it from T. buettneri include asymmetrical, tricuspate teeth in which the central cusp is taller than the marginal cusp, polygonal in cross-section and displaced slightly lingually; the lingual cusp is low, transversely broadened and anteroposteriorly compressed; teeth with prominent cingula along the mesial and distal margins; and paired sagittal (parasagittal) crests composed of portions of the postfrontals and the parietals, as well as a relatively broad skull roof between the supratemporal fenestrae. We also re-illustrate and re-describe the holotypes of both Trilophosaurus buettneri Case and T. jacobsi Murry. T. jacobsi is primarily Adamanian in age, and the lowest occurrence of T. jacobsi is stratigraphically above that of T. buettneri. Therefore, T. buettneri is an Otischalkian-Adamanian index taxon, and T. jacobsi is an Adamanian-?earliest Revueltian index taxon. Trilophosaurus teeth are readily identifiable to species and therefore identifiable as isolated fossils.

Located in Santa Fe County, New Mexico, USA, the Lamy amphibian quarry is a Late Triassic (Adamanian) bonebed stratigraphically low in the Garita Creek Formation of the Chinle Group. Well known for its mass accumulation of metoposaurid amphibians, it was initially interpreted as a drought-induced death assemblage. Based on microstratigraphic and sedimentological studies, additional and extensive collecting at the quarry and a revised understanding of the bonebed, we provide a detailed taphonomic analysis of the Lamy amphibian quarry that identifies it as a low diversity multitaxic and monodominant bonebed in pedogenically modified floodplain mudstone. The Lamy bonebed shows no evidence of drought and is characterized by a high density of completely dissociated bones that show clear alignment by current and sorting (enrichment of Voorhies Group II and III elements). The bones show no significant abrasion or weathering (stage 0), preserve virtually no evidence of scavenging and show no evidence of trampling. Based on skull lengths, the metoposaurid assemblage has a type I survivorship curve and lacks juveniles. We thus posit that the following sequence of events formed the Lamy amphibian bonebed: (1) aggregation (cause unknown) of a large number of metoposaurid amphibians at a site different from the location of the bonebed, though not distant; (2) catastrophic mass mortality; (3) complete disarticulation and disassociation of the skeletons; and (4) rapid transport of the disarticulated bones onto a floodplain surface that was undergoing pedogenesis. The Lamy amphibian bonebed is representative of the Late Triassic metoposaurid bonebeds from Morocco and the western USA, which are monodominant and nearly monotaxic. They indicate that aggregation (probably of breeding populations) and mass death of metoposaurids were relatively common across the riverine floodplains of Late Triassic Pangea. Fig. 9. Schematic representation of events that lead to the formation of the Lamy amphibian bonebed. A population of metoposaurs died at one location, decayed to disarticulated bones, and these bones were transported to the current location of the Lamy amphibian bonebed.

Doswellia sixmilensis is a new species of the doswelliid archosauromorph genus Doswellia named for an incomplete skeleton from the Upper Triassic Bluewater Creek Formation of the Chinle Group in west‐central New Mexico, USA. D. sixmilensis differs from D. kaltenbachi Weems, the type and only other known species of Doswellia, in its larger size, higher tooth count and greater heterodonty, possession of keels on the cervical centra and the presence of discrete knobs or spikes on some osteoderms. The holotype of D. sixmilensis is the fourth occurrence of Doswellia and only the second occurrence of a Doswellia skull, which includes the previously unknown premaxilla and maxilla (and therefore the best dentition) and has the best‐preserved cervical vertebrae. Although it adds to our knowledge of the anatomy of Doswellia, this new information does not alter previous concepts of the phylogenetic relationships of the doswelliid genera, largely because they are so poorly known anatomically. The genus Doswellia is known from the Newark Supergroup in Virginia, and the Chinle Group in Texas, New Mexico and Utah, in strata of Otischalkian–Adamanian age. The type locality of D. sixmilensis is c. 43 m stratigraphically below a bed from which U‐Pb dating of detrital zircons yields a maximum depositional age of c. 220 Ma, so this is a reasonable approximate numerical age for D. sixmilensis.