mains, with loss increasing by 210 100 ha yr −1 (Hansen et al. 2013). Between 1990 and 2010, tropical forest cover was reduced from 1635 mil lion ha to 1514 million ha, with 32.9 million ha lost in Southeast Asia (Achard et al. 2014). Overexploitation and agricultural activities are the primary threats to species worldwide

The space in which organisms live determines health and physicality, shaping the way in which they interact with their peers. Space, therefore, is critically important for species diversity and the function performed by individuals within mixed communities. The biotic and abiotic factors defined by the space that organisms occupy are ecologically significant and the difficulty in quantifying space-defined parameters within complex systems limits the study of ecological processes. Here, we overcome this problem using a tractable system whereby spatial heterogeneity in interacting fungal wood decay communities demonstrates that scale and patchiness of territory directly influence coexistence dynamics. Spatial arrangement in 2-and 3dimensions resulted in measurable metabolic differences that provide evidence of a clear biological response to changing landscape architecture. This is of vital importance to microbial systems in all ecosystems globally, as our results demonstrate that community function is driven by the effects of spatial dynamics.