). The Z/K ratio (Z/K = 2.65 for S. mandibularis; Z/K = 4.85 for S. intermedius) indicated that mortality was predominant over growth for the two species. The recruitment pattern showed one Gauss curve translates was continuous for each species. For S. mandibularis, the exploitation rate (E = 0.47) was close to E 0.1 (E 0.1 = 0.46), indicating that S. mandibularis's stock was in an optimum state of exploitation. However, for S. intermedius, E max was lower (E max = 0.57) than the exploitation rate (E = 0.77). This result reflected overexploitation of this species.
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