Abstract. Syngenetic permafrost deposits formed extensively on and around the arising Beringian subcontinent during the Late Pleistocene sea level lowstands. Syngenetic deposition implies that all material, both mineral and organic, freezes parallel to sedimentation and remains frozen until degradation of the permafrost. Permafrost is therefore a unique archive of Late Pleistocene palaeoclimate. Most studied permafrost outcrops are situated in the coastal lowlands of northeastern Siberia; inland sections are, however, scarcely available. Here, we describe the stratigraphical, cryolithological, and geochronological characteristics of a permafrost sequence near Batagay in the Siberian Yana Highlands, the interior of the Sakha Republic (Yakutia), Russia, with focus on the Late Pleistocene Yedoma ice complex (YIC). The recently formed Batagay mega-thaw slump exposes permafrost deposits to a depth of up to 80 m and gives insight into a climate record close to Verkhoyansk, which has the most severe continental climate in the Northern Hemisphere. Geochronological dating (optically stimulated luminescence, OSL, and 14C ages) and stratigraphic implications delivered a temporal frame from the Middle Pleistocene to the Holocene for our sedimentological interpretations and also revealed interruptions in the deposition. The sequence of lithological units indicates a succession of several distinct climate phases: a Middle Pleistocene ice complex indicates cold stage climate. Then, ice wedge growth stopped due to highly increased sedimentation rates and eventually a rise in temperature. Full interglacial climate conditions existed during accumulation of an organic-rich layer – plant macrofossils reflected open forest vegetation existing under dry conditions during Marine Isotope Stage (MIS) 5e. The Late Pleistocene YIC (MIS 4–MIS 2) suggests severe cold-stage climate conditions. No alas deposits, potentially indicating thermokarst processes, were detected at the site. A detailed comparison of the permafrost deposits exposed in the Batagay thaw slump with well-studied permafrost sequences, both coastal and inland, is made to highlight common features and differences in their formation processes and palaeoclimatic histories. Fluvial and lacustrine influence is temporarily common in the majority of permafrost exposures, but has to be excluded for the Batagay sequence. We interpret the characteristics of permafrost deposits at this location as a result of various climatically induced processes that are partly seasonally controlled. Nival deposition might have been dominant during winter time, whereas proluvial and aeolian deposition could have prevailed during the snowmelt period and the dry summer season.

Abstract. Ice wedges in the Yana Highlands of interior Yakutia – the most continental region of the Northern Hemisphere – were investigated to elucidate changes in winter climate and continentality that have taken place since the Middle Pleistocene. The Batagay megaslump exposes ice wedges and composite wedges that were sampled from three cryostratigraphic units: the lower ice complex of likely pre-Marine Isotope Stage (MIS) 6 age, the upper ice complex (Yedoma) and the upper sand unit (both MIS 3 to 2). A terrace of the nearby Adycha River provides a Late Holocene (MIS 1) ice wedge that serves as a modern reference for interpretation. The stable-isotope composition of ice wedges in the MIS 3 upper ice complex at Batagay is more depleted (mean δ18O about −35 ‰) than those from 17 other ice-wedge study sites across coastal and central Yakutia. This observation points to lower winter temperatures and therefore higher continentality in the Yana Highlands during MIS 3. Likewise, more depleted isotope values are found in Holocene wedge ice (mean δ18O about −29 ‰) compared to other sites in Yakutia. Ice-wedge isotopic signatures of the lower ice complex (mean δ18O about −33 ‰) and of the MIS 3–2 upper sand unit (mean δ18O from about −33 ‰ to −30 ‰) are less distinctive regionally. The latter unit preserves traces of fast formation in rapidly accumulating sand sheets and of post-depositional isotopic fractionation.

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Megafaunal extinctions are recurring events that cause evolutionary ripples, as cascades of secondary extinctions and shifting selective pressures reshape ecosystems. Megafaunal browsers and grazers are major ecosystem engineers, they: keep woody vegetation suppressed; are nitrogen cyclers; and serve as seed dispersers. Most angiosperms possess sets of physiological traits that allow for the fixation of mutualisms with megafauna; some of these traits appear to serve as exaptation (preadaptation) features for farming. As an easily recognized example, fleshy fruits are, an exaptation to agriculture, as they evolved to recruit a non-human disperser. We hypothesize that the traits of rapid annual growth, self-compatibility, heavy investment in reproduction, high plasticity (wide reaction norms), and rapid evolvability were part of an adaptive syndrome for megafaunal seed dispersal. We review the evolutionary importance that megafauna had for crop and weed progenitors and discuss possible ramifications of their extinction on: (1) seed dispersal; (2) population dynamics; and (3) habitat loss. Humans replaced some of the ecological services that had been lost as a result of late Quaternary extinctions and drove rapid evolutionary change resulting in domestication.