Phenology is changing in response to anthropogenic climate change (Parmesan and Yohe, 2003) and understanding exactly how phenological shifts track changing environmental conditions can help prioritize mitigation as well as management actions (Morellato et al., 2016). For example, if we can predict the phenology of recently arrived invasive species based on their closest relatives, we can narrow down treatment windows for manual removal or herbicide application.Phenological studies of flowering plants use combinations of historical observations and herbarium specimens to track changes in leaf, floral, and fruiting phenology (Primack et al., 2004;Munson and Long, 2017;Willis et al., 2017;Meineke et al., 2019). The multiple lifetimes of collection efforts housed in herbaria span space and time, and provide irreplaceable insight into long-term trends and broadscale patterns of changes in phenology (
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