SummaryThe mitochondrial DNA variation of 295 Berber-speakers from Morocco (Asni, Bouhria and Figuig) and the Egyptian oasis of Siwa was evaluated by sequencing a portion of the control region (including HVS-I and part of HVS-II) and surveying haplogroup-specific coding region markers. Our findings show that the Berber mitochondrial pool is characterized by an overall high frequency of Western Eurasian haplogroups, a somehow lower frequency of sub-Saharan L lineages, and a significant (but differential) presence of North African haplogroups U6 and M1, thus occupying an intermediate position between European and sub-Saharan populations in PCA analysis. A clear and significant genetic differentiation between the Berbers from Maghreb and Egyptian Berbers was also observed. The first are related to European populations as shown by haplogroup H1 and V frequencies, whereas the latter share more affinities with East African and Nile Valley populations as indicated by the high frequency of M1 and the presence of L0a1, L3i, L4 * , and L4b2 lineages. Moreover, haplogroup U6 was not observed in Siwa. We conclude that the origins and maternal diversity of Berber populations are old and complex, and these communities bear genetic characteristics resulting from various events of gene flow with surrounding and migrating populations.

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In the typology of morphological borrowing, one type has received little attention: cases where words are borrowed in several paradigmatic forms. An example of this is found in English alumnus -alumni, where Latin nouns are borrowed both in their singular and plural forms. Such borrowings lead to a coexistence of borrowed and native paradigms in one and the same language. This type of borrowing is called Parallel System Borrowing (PSB). Such patterns are wide-spread, and concern virtually all parts of morphology, including verbal inflection and pronouns. The emergence of PSB is not governed by a single sociolinguistic factor, such as the existence of learned registers (as with alumnus -alumni). In fact, it appears that some of the most spectacular cases of PSB have no relation to learned registers at all.

Several verbal forms reconstructed for proto-Semitic strongly resemble reconstructed forms in proto-Berber: compare Semitic yV-PaRRaS to Berber y-əFăRRăS, Semitic yV-PRaS to Berber y-əFRăS, and Semitic yV-PRuS and yV-PRiS to Berber y-ăFRəS. We suggest that these forms are historically related and sketch a line of development from the reconstructed meanings to their attested uses. yVPaRRaS, originally imperfective, retains that value in both Berber and Semitic. yVPRas, originally stative, gained a perfective meaning in Berber and Semitic; the stative meaning is retained in Berber, but was largely lost in Semitic. yVPRus/yVPRiS, originally perfective, retained that meaning in Semitic, merging with the newly perfective yVPRas forms; in Berber, yVPRaS completely replaced perfective yVPRuS/yVPRiS, relegating the latter to non-aspectual uses. We conclude by considering the quality of the first vowel; the alternation seen in Berber y-əFRăS and y-ăFRəS supports reconstructions as yiPRaS and yaPRuS/yaPRiS, conforming to the Barth–Ginsberg Law of Semitic.

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