Abstract. Water levels in streams and aquifers often exhibit daily cycles during rainless periods, reflecting daytime extraction of shallow groundwater by evapotranspiration (ET) and, during snowmelt, daytime additions of meltwater. These cycles can aid in understanding the mechanisms that couple solar forcing of ET and snowmelt to changes in streamflow. Here we analyze 3 years of 30 min solar flux, sap flow, stream stage, and groundwater level measurements at Sagehen Creek and Independence Creek, two snow-dominated headwater catchments in California's Sierra Nevada mountains. Despite their sharply contrasting geological settings (most of the Independence basin is glacially scoured granodiorite, whereas Sagehen is underlain by hundreds of meters of volcanic and volcaniclastic deposits that host an extensive groundwater aquifer), both streams respond similarly to snowmelt and ET forcing. During snow-free summer periods, daily cycles in solar flux are tightly correlated with variations in sap flow, and with the rates of water level rise and fall in streams and riparian aquifers. During these periods, stream stages and riparian groundwater levels decline during the day and rebound at night. These cycles are reversed during snowmelt, with stream stages and riparian groundwater levels rising during the day in response to snowmelt inputs and falling at night as the riparian aquifer drains. Streamflow and groundwater maxima and minima (during snowmelt- and ET-dominated periods, respectively) lag the midday peak in solar flux by several hours. A simple conceptual model explains this lag: streamflows depend on riparian aquifer water levels, which integrate snowmelt inputs and ET losses over time, and thus will be phase-shifted relative to the peaks in snowmelt and evapotranspiration rates. Thus, although the lag between solar forcing and water level cycles is often interpreted as a travel-time lag, our analysis shows that it is mostly a dynamical phase lag, at least in small catchments. Furthermore, although daily cycles in streamflow have often been used to estimate ET fluxes, our simple conceptual model demonstrates that this is infeasible unless the response time of the riparian aquifer can be determined. As the snowmelt season progresses, snowmelt forcing of groundwater and streamflow weakens and evapotranspiration forcing strengthens. The relative dominance of snowmelt vs. ET can be quantified by the diel cycle index, which measures the correlation between the solar flux and the rate of rise or fall in streamflow or groundwater. When the snowpack melts out at an individual location, the local groundwater shifts abruptly from snowmelt-dominated cycles to ET-dominated cycles. Melt-out and the corresponding shift in the diel cycle index occur earlier at lower altitudes and on south-facing slopes, and streamflow integrates these transitions over the drainage network. Thus the diel cycle index in streamflow shifts gradually, beginning when the snowpack melts out near the gauging station and ending, months later, when the snowpack melts out at the top of the basin and the entire drainage network becomes dominated by ET cycles. During this long transition, snowmelt signals generated in the upper basin are gradually overprinted by ET signals generated lower down in the basin. The gradual springtime transition in the diel cycle index is mirrored in sequences of Landsat images showing the springtime retreat of the snowpack to higher elevations and the corresponding advance of photosynthetic activity across the basin. Trends in the catchment-averaged MODIS enhanced vegetation index (EVI2) also correlate closely with the late springtime shift from snowmelt to ET cycles and with the autumn shift back toward snowmelt cycles. Seasonal changes in streamflow cycles therefore reflect catchment-scale shifts in snowpack and vegetation activity that can be seen from Earth orbit. The data and analyses presented here illustrate how streams can act as mirrors of the landscape, integrating physical and ecohydrological signals across their contributing drainage networks.

Measurements of freshwater mussel tissue are potentially very useful for determining base-level isotopic values for food web studies in aquatic environments. As long-lived, filter-feeding organisms, mussels have the potential to spatially and temporally average the isotopic baseline signal. Following from earlier studies that focused on lake environments, this study investigates the stable carbon and nitrogen isotope ratios in tissue of the river dwelling freshwater mussel, Margaritifera falcata, in two extensively studied northern California coast range rivers, the South Fork Eel and Navarro. We highlight advantages and challenges for using riverine mussel isotopes as indicators of baselines. d 13 C of primary producers is known to vary with habitat along the South Fork Eel channel, but our measurements show no such variations, demonstrating that riverine mussels do preserve a spatially averaged measure of instream derived food sources. Mean d 13 C and d 15 N are shown to be markedly different in the two rivers, reflecting differences in food sources and possibly watershed land use. We also found that d 15 N of mussel tissue increased by approximately 2& with mussel age in both rivers. This suggests it is important to consider age and size effects when estimating baseline values from mussel tissues.