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Few of Stephen Jay Gould's accomplishments in evolutionary biology have received more attention than his hierarchical theory of evolution, which postulates a causal discontinuity between micro- and macroevolutionary events. But Gould's hierarchical theory was his second attempt to supply a theoretical framework for macroevolutionary studies-and one he did not inaugurate until the mid-1970s. In this paper, I examine Gould's first attempt: a proposed fusion of theoretical morphology, multivariate biometry and the experimental study of adaptation in fossils. This early "macroevolutionary synthesis" was predicated on the notion that parallelism and convergence dominate the history of higher taxa, and moreover, that they can be explained in terms of adaptation leading to mechanical improvement. In this paper, I explore the origins and contents of Gould's first macroevolutionary synthesis, as well as the reasons for its downfall. In addition, I consider how various developments during the mid-1970s led Gould to identify hierarchy and constraint as the leading themes of macroevolutionary studies-and adaptation as a macroevolutionary red herring.

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This article is about a start-up problem in scientific practice. Specifically, it is about the problem of justifying paleontological correlation—the practice of using fossils to establish time relations among fossiliferous rocks. Paleontological correlation was the key to assembling a geological timescale during the nineteenth century and remains an important practice in stratigraphic geology to this day. Yet contrary to philosophical expectations, this practice lacked a robust theoretical justification during the first half of the nineteenth century. This article examines what this lack of justification amounted to, as well as how the method of paleontological correlation was justified in practice.

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