Microbialite masses within the lowest Triassic strata along the southern periphery of the tropical Yangtze Platform were produced in a post-Permian microbial regime. These microbialites (the Hindeodus parvus Zone), which are represented by thrombolites, occur only where terrigenous sediment influx was rare, even in shallow-marine settings. The lower parts of the thrombolites lie upon a distinctly unconformable Permian-Triassic boundary and exhibit a thinbedded to thick-bedded planar structure. In contrast, the upper parts of the thrombolites contain domed macrostructures that interact in complex ways with skeletal grainstones and packstones. Irregular frameworks of thrombolite bodies differ in degree of lateral and vertical accumulation and in the amalgamation of mesoclots of microbial origin; they exhibit marked variations in texture. A transgressive episode occurred in the earliest Triassic following the mass extinctions, and this included the initiation of microbial regimes that usually formed planar thrombolite masses in lower-energy, deep subtidal environments. The varied textures and structures of thrombolites during deposition may reflect a combination of sea-level fluctuations, physicochemical ocean conditions, microbial activity, skeletalsediment influx, and other factors. These earliest Triassic, uniquely microbial regimes collapsed in stepwise fashion and were succeeded by the Isarcicella staeschei and I. isarcica zones, which contain a predominance of mudstones, suggesting a marked sea-level transgression. Space-specific and time-specific, the earliest Triassic microbialites record short-term, high-resolution paleoenvironmental fluctuations immediately after the end-Permian extinctions.
The Ordovician is a period when novel reef ecosystems appeared along with new reef constructors and skeletal-dominated reefs. The Lower Ordovician (late Tremadocian) Fenhsiang Formation of the Three Gorges area in South China contains the oldest known bryozoan reefs (lithistid sponge-bryozoan and bryozoan-pelmatozoan reefs) alongside lithistid sponge-microbial reefs. The latter are characterized by the dominance of microbialites that encrusted and bound the frame-building sponges and inter-sponge sediments. In contrast, the lithistid sponge-bryozoan and bryozoan-pelmatozoan reefs are generally characterized by bryozoans that encrusted the frame-building sponges or pelmatozoans and grew to fill the inter-frameworks. These sponges and pelmatozoans did not construct the rigid frameworks unaided; their association with bryozoans enabled the development of small skeletal-dominated reefs with rigid frameworks. Skeletal-dominated reefs, for which frame-constructing and encrusting roles are conspicuous, were largely unknown before the Early Ordovician. The appearance of skeletal organisms (specifically colonial, encrusting bryozoans) enabled the development of skeletal-dominated reefs, which were pioneers in the rise of Middle-Late Ordovician reefs. The Early Ordovician establishment of skeletal-dominated reefs at the earliest stages of the Great Ordovician Biodiversification Event would have created novel niches and biological interactions that further promoted the evolution of reef-building and -dwelling organisms, as well as ensuing reef ecosystems.
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