Dromedaries have been fundamental to the development of human societies in arid landscapes and for long-distance trade across hostile hot terrains for 3,000 y. Today they continue to be an important livestock resource in marginal agro-ecological zones. However, the history of dromedary domestication and the influence of ancient trading networks on their genetic structure have remained elusive. We combined ancient DNA sequences of wild and early-domesticated dromedary samples from arid regions with nuclear microsatellite and mitochondrial genotype information from 1,083 extant animals collected across the species' range. We observe little phylogeographic signal in the modern population, indicative of extensive gene flow and virtually affecting all regions except East Africa, where dromedary populations have remained relatively isolated. In agreement with archaeological findings, we identify wild dromedaries from the southeast Arabian Peninsula among the founders of the domestic dromedary gene pool. Approximate Bayesian computations further support the "restocking from the wild" hypothesis, with an initial domestication followed by introgression from individuals from wild, now-extinct populations. Compared with other livestock, which show a long history of gene flow with their wild ancestors, we find a high initial diversity relative to the native distribution of the wild ancestor on the Arabian Peninsula and to the brief coexistence of early-domesticated and wild individuals. This study also demonstrates the potential to retrieve ancient DNA sequences from osseous remains excavated in hot and dry desert environments.anthropogenic admixture | Camelus dromedarius | demographic history | paleogenetics | wild dromedary T he dromedary (Camelus dromedarius) is one of the largest domestic ungulates and one of the most recent additions to livestock. Known as the "ship of the desert" (1), it enabled the transportation of people and valuable goods (e.g., salt, incense, spices) over long distances connecting Arabia, the Near East, and North Africa. This multipurpose animal has outperformed all other domestic mammals, including the donkey, in arid environments and continues to provide basic commodities to millions of people inhabiting marginal agro-ecological zones. In the current context of advancing desertification and global climate change, there is renewed interest in the biology and production traits of the species (2), with the first annotated genome drafts having been recently released (3, 4). SignificanceThe dromedary is one of the largest domesticates, sustainably used in arid and hostile environments. It provides food and transport to millions of people in marginal agricultural areas. We show how important long-distance and back-and-forth movements in ancient caravan routes shaped the species' genetic diversity. Using a global sample set and ancient mitochondrial DNA analyses, we describe the population structure in modern dromedaries and their wild extinct ancestors. Phylogenetic analyses of ancient and modern dro...

The archaeology of prehistoric Arabia : adaptation and social formation from the neolithic to the iron age / Peter Magee. pages cm.-(Cambridge world archaeology) Includes bibliographical references and index. ISBN 978-0-521-86231-8 (hardback) 1. Prehistoric peoples-Arabian Peninsula. 2. Excavations (Archaeology)-Arabian Peninsula. 3. Social archaeology-Arabian Peninsula. 4. Arabian Peninsula-Antiquities. I. Title. GN855.A72M25 2014 939.4′9-dc23 2013027340 ISBN 978-0-521-86231-8 Hardback Cambridge University Press has no responsibility for the persistence or accuracy of URLs for external or third-party Internet Web sites referred to in this publication and does not guarantee that any content on such Web sites is, or will remain, accurate or appropriate.

Drawing on the results of the Australian Archaeological Expedition's excavation of Tell Abraq, a three‐fold division of the southeast Arabian Iron Age is suggested. A re‐analysis of 14C data in line with the latest agreed calibration curve together with an analysis of the foreign parallels of southeast Arabian Iron Age pottery permits the construction of a complete Iron Age sequence. The results argue for a lower chronology for the Iron II and III periods than has, up until now, been suggested.

This article summarizes the outcome of a workshop sponsored by the Durham University Centre for Iranian Cultural Studies, where papers were presented on the entire chronological range of water management systems in Iran from around 8000 years bc until around 1000 ad. The primary aim was to recognize major research questions that could be used to create an agenda for future studies of ancient water use in the country. In the Durham meeting, it appeared that although the small-scale prehistoric systems probably constituted an example of ‘human niche construction’, the later imperial systems did not. Despite the recognition of occasional irrigation systems of third millennium bc date in the Deh Luran plain by Neely and Wright, as well as perhaps in Khuzestan, there appears to be a general dearth of evidence of Chalcolithic and Bronze Age systems in Iran. However, by the first millennium bc there was a considerable increase in the construction of major water management systems, some of which were, at least as far as the associated evidence suggests, constructed by imperial authorities. All agreed, however, that just because a system appeared large in scale, it was not necessarily a result of imperial management. For the subject of qanats it was argued that not only were they usually built by small-scale societies, but also that there may have been multiple centres of origin; one primary centre being a broad zone of south-east Iran, Pakistan and south-east Arabia

Research recommenced at Tell Abraq in 2007, carried out by Bryn Mawr College with an international team in co-operation with the Sharjah Archaeology Authority. Between 2007 and 2010 fieldwork was limited to documenting sections of the previous excavations and one small sounding while our attention focused mainly on the nearby shell-midden site of Hamriya and the fortified Iron Age settlement site of Muweilah. Since 2010, large-scale excavations have been conducted which have fundamentally altered our understanding of the layout of the settlement and its economic relations during the second and first millennia BC. In this report, we detail the results of this work and present preliminary interpretations. K E Y W O R D S Bronze Age, dromedary domestication, Iron Age, settlement mound, south-east Arabia F I G U R E 1 A map showing the location of Tell Abraq and other sites mentioned in the text