The thorax of Hypodicranotus has ten segments and a spine on the eighth. The ages of Erratencrinurus s.l. spicatus and Erratencrinurus (Erratencrinurus?) vigilans in the Lake St. John district do not confirm their temporal roles leading to subgenera of Erratencrinurus, as has been recently suggested. Phylogenetic analyses of large data sets of species previously referred to Encrinuroides and Physemataspis yield a minimal length cladogram containing 18 species. Encrinuroides is restricted to four species, two of which have biogeographic affinities with Iapetus. These results lead to three clades, named the Walencrinuroides n. gen. clade, Frencrinuroides n. gen. clade, and finally the Physemataspis clade, with an enlarged concept of the genus with the erection of Physemataspis (Prophysemataspis) n. subgen. These last three clades are restricted to North America and Scotland, with alternating predominance of one region. Walencrinuroides s.l. gelaisi n. gen. n. sp. is described. New morphological data on Erratencrinurus s.l. spicatus confirm its close relationship with the clades discussed above. Data are insufficient for phylogenetic analysis of selected cheirurine species here surveyed. Eye position, glabellar segmentation, and pygidial shape differentiate the genera Ceraurus and Gabriceraurus; emended diagnoses of these genera are presented. Ceraurus globulobatus and C. matranseris are distinct, but morphologically close to one another. The status of Gabriceraurus dentatus can be stabilized on its extant types.
The Ordovician–Silurian strata of the Percé area are assigned to the Matapédia Group. They occur in a southwestern monoclinal sequence, unconformably overlying Cambrian strata, and are assigned to the Pabos and White Head formations. The Pabos Formation is preponderantly a terrigenous sequence, whereas the White Head is preponderantly a carbonate sequence. The Pabos strata of this area are included in the new Rouge Member, in which four brachiopod-dominated communities are recognized: the Dalmanella, Catazyga, Sowerbyella, and Epitomyonia communities, within which trilobites occur sporadically. The White Head Formation is divided into three new limestone members and a new mudstone member. The basal Burmingham Member has yielded a Catazyga Community and a Sowerbyella-like Community. The Côte de la Surprise Member is composed of mudstones with the previously described Hirnantia Community. The Rouge, Burmingham, and Côte de la Surprise members are Ashgillian (Upper Ordovician), but the uppermost two members of the White Head Formation, the L'Irlande and Des Jean members, are Llandoverian. They yield an Acernaspis Community, assigned a paleoecological position intermediate between those of the Clorinda and graptolite communities. The Matapédia Group limestones and shales in the structurally complex northeastern sequence are informally termed the Grande Coupe beds. These beds are partly or wholly time-correlative with the Rouge and Burmingham members but were deposited in deeper water. A Stenopareia Community includes the highly fossiliferous Grande Coupe beds, with a local development of the Foliomena Community. The Percé area is unique within the Quebec Appalachians because the strata of the Matapédia Group are highly fossiliferous, with distinct European affinities in the Ordovician, and because the monoclinal sequence is a deepening-upward sequence, probably to the north and west of deeper water clastics.
The Ashgillian Pabos Formation underlies the Ashgillian-Llandoverian White Head Formation and consists, in the immediate Percé area, of 21 trilobite taxa. Of these, 11 are left in open nomenclature, four of which are summarily described, commented upon, and illustrated. Ten species are specifically determined and three of these are new: Remopleurides arenosus, Achatella (Achatella) clivosa, and Meadowtownella sacerdos. Hibbertia whittingtoni (McNamara, 1979) is a junior secondary homonym and renamed H. conistonensis. The Ashgillian-Llandoverian cheirurines of the Percé area are reviewed and new generic assignments suggested; these are a major component of the Pabos trilobite fauna. Revision of Ceraurinus icarus, a widely distributed North American species, shows its inherent variability; limits of variation, more complete descriptions, and stratigraphic distribution of most previously described species are also given. The distinctive trinucleid Novaspis elevata occurs in the Pabos, outside the immediate Perce area. Almost all of the Pabos trilobites are also present in the coeval Grande Coupe beds of the Matapedia Group; otherwise they are a mixture of taxa with greatest faunal affinities with North American or northwestern European taxa of the same age.
Four previously defined formations within the Chicoutimi outlier are extended to the Lac Saint-Jean outlier; these formations are gradational into one another. The lowest of these formations, the Tremblay, is restricted to siliciclastic strata (predominantly coarse-grained sandstones); the overlying Simard, to micritic limestones. This last-named formation is succeeded by the highly fossiliferous Shipshaw Formation, composed of alternating limestones and shales, whereas the youngest of the previously named carbonates, the Galets Formation, is made up of echinodermal calcarenites. Graptolitic shales, herein assigned to the new Pointe-Bleue Shales, overlie in sharp contact the Galets; the youngest limestones of the area, previously assigned to the Richmondian, are different from the other Ordovician limestones. Trilobite faunas from the Shipshaw (and Galets) are correlated with the Edenian; the same correlation is probable for the uppermost of the four informal units recognized within the Simard Formation. The lowest three informal units of the Simard Formation are assigned to the Kirkfieldian; Shermanian strata have not been identified but are probably present. The Pointe-Bleue Shales were previously assigned to the Maysvillian. The Simard Formation contains an "Arctic" fauna, also present in the Manicouagan outlier but yielding in this last case a Shipshaw–Galets trilobite fauna. The Simard fauna, previously correlated with the "Black River," is thus interpreted as significantly time transgressive and of limited stratigraphic use. Depositional environments present in the Lac Saint-Jean and Chicoutimi outliers, but absent in southern Ontario during the Shermanian and Edenian, account for the necessity of significantly increasing the biozones of numerous taxa.
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