Three wheat experiments are described in which a range of plant populations were shaded during different periods of development; in two of the experiments plant thinning was also carried out at a number of growth stages. Shading during the period of ear development caused an appreciable decrease in grain yield by decreasing the number of grains per ear. Shading during the grain filling period also reduced grain yield, this being brought about by decreased grain size. Thus in contrast to the barley experiments reported earlier (Willey & Holhday, 1971), these particular results gave no indication of a potential surplus of carbohydrate for grain filling and an associated limited ear capacity. However, when plant thinning was carried out at anthesis to make more carbohydrate available for grain filling in the remaining ears, grain yield per ear did not increase. It is argued, therefore, that grain yield probably was determined at least partly by a limited ear capacity. Plant thinning at earlier stages showed how the development of competition during the ear development period progressively reduced the potential capacity of the ear; the greater competition of higher plant populations accelerated this reduction in ear potential.From an examination of the effects of plant population, it is suggested that the number of grains per ear is the component having greatest influence on the decline in grain yield at above-optimum populations. The possible importance of the number of grains per unit area as an indicator of ear capacity on an area basis, and as a determinant of grain yield per unit area, is emphasized. A close relationship between grain yield per unit area and number of grains per unit area is illustrated for a number of plant-population response curves, and it is suggested that the decrease in grain yield at high populations is probably determined by a decrease in the number of grains per unit area. Evidence is presented to substantiate the idea put forward in the barley paper that this decrease in the number of grains per unit area may be attributable more to a lower production of total dry matter by the high populations during the later stages of ear development, than to an unfavourable partitioning of such dry matter between the ear and the rest of the plant. INTRODUCTIONt o t h a t U 8 e d m t h e b a r l e v m t n a t a r a n 8 e o f plan* populations were shaded during different periods An earlier paper (Willey & Holliday, 1971) of development. In two of the experiments the reported the effects of plant population and shading range of plant populations was much greater than in some barley experiments. This paper deals with that used in the barley and a more detailed examinathree similar wheat experiments. The main purpose tion of the various development periods was underwas to examine the physiological factors deter-taken. Also, in these same two experiments, selected mining the relationship between plant population treatments were thinned at different stages of and grain yield, particularly the decrease in...
The effects of 4 levels of applied nitrogen, ranging from nil to a maximum of 417 lb N/ acre/annum, in all combinations with 3 frequencies of defoliation, ranging from 2 to a maximum of 10 cuts per annum, on herbage production from a perennial ryegrass/ timothy/meadow fescue/white clover sward were measured. These treatments were operative for 2J years, and in a subsequent year the residual effect of cutting frequency was tested. Dry-matter yields of total herbage and of the clover fraction are quoted, together with N yields of total herbage. Yield response to N was higher than in some other experiments in the U.K. Cutting frequency had a very large effect and, in general, the longer the interval between cuts, the higher was the dry-matter (though not the N) yield. There was a marked interaction between cutting frequency and level of N: at the high cutting frequency, dry-matter yield increased linearly with increasing level of N; at the medium frequency, response tended to fall off at the highest level of N; at the low frequency, yield declined with increasing level of N beyond 139 lb N per acre per anum.
The objective of this study was to explore, using first-hand accounts, adolescents' understandings of why they self-harmed, what their responses to self-harm were, and how they resisted or ceased self-harm. Secondary analysis was conducted of video-recorded family therapy sessions from the Self-harm Intervention: Family Therapy (SHIFT Trial). Recordings of 22 participants, approximately 170 hours of footage, formed the dataset. The study developed 5 core themes: (1) Distress can be difficult to convey; (2) Self-harm and suicidal ideation: a complex relationship; (3) Self-harm as a form of communication; (4) Self-harm to manage emotions; and (5) Moving forward. Self-harm was a means of communicating distress as well as managing emotions. Accounts highlighted the complex interplay between self-harm and suicidal intent. Encouragingly, many participants described being able to resist self-harm.
Two barley experiments are described in which a range of plant populations were shaded during different periods of development. Shading during the ear development period caused considerable reductions in grain yield, largely by reducing the number of grains per ear. Shading during the grain-filling period caused no reduction in grain yield. It is suggested that under conditions of these experiments there was probably a potential surplus of carbohydrate available for grain filling and that grain yield was largely determined by the storage capacity of the ears. The importance of the number of grains per ear as an indicator of individual ear capacity is emphasized.The effects of plant population on grain yield and its components are also examined. It is concluded that the number of grains per ear is the component having greatest influence on the decrease in grain yield at above-optimum populations and attention is again drawn to the possible importance of ear capacity. It is argued that on an area basis the number of grains per unit area may give a good indication of ear capacity. Examination of this parameter shows a close relationship with grain yield per unit area for both the shading and population treatments. It is particularly evident that a decrease in grain yield at high populations was associated with a comparable decrease in the number of grains per unit area. It is suggested that this decrease in grain number may be due to a lower production of total dry matter during ear development rather than an unfavourable partitioning of this dry matter between the ear and the rest of the plant. This lower production of total dry matter is attributed to the crop growth rates of the higher populations having reached their peak and then having declined before the end of the ear development period. This crop growth rate pattern, through its effect on grain number per unit area, is put forward as the basic reason why, in the final crop, grain yield per unit area decreases at above-optimum populations.
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