The Asteropyginae Delo, 1935 is a group of phacopid trilobites in the family Acastidae Delo, 1935 that has served as the focus for several studies due to their distinctive morphologies and diversity. However, despite an interest in these characteristic morphologies, there have been no studies that have examined this group using morphometric techniques. Our investigation utilized both geometric morphometric and elliptical Fourier methods to quantify the morphology of cephalic sclerites of asteropyginid specimens representing wide taxonomic sampling of the clade. We constructed a phylomorphospace that shows temporal and spatial patterns of phenotypic evolution within the framework of a novel tip-dated phylogenetic tree generated using Bayesian inference. We recovered similar patterns in disparity regardless of the morphometric approach. Both analyses illustrated a marked expansion into morphospace throughout the temporal range of the clade, peaking in disparity in the Emsian and with European taxa exhibiting the highest disparity in glabellar morphospace. Additionally, glabellar shape showed low phylogenetic signal and no major patterns in phylomorphospace. This study highlights the utility of employing different methodologies to quantitatively explore the disparity of fossil taxa. It also illustrates some of the patterns of morphological change occurring during one of the final and major evolutionary radiations within Phacopida.
Ontogenetic sequencing of suture fractal dimensions from a single ammonite specimen was not previously accessible without extreme measures. The necessity for destruction of the full conch due to whorl overlap or, alternatively, the prohibitively expensive or inaccessible imaging equipment for use by paleontologists has led to this rich source of data remaining elusive. Tracings of ontogenetic sequences of sutures have rarely been published since the middle of the last century. These studies only focused on the outermost whorl of a given specimen or composited several specimens. Oftentimes, they focused only on the umbilical and juvenile stages of shell growth. Complete or multi-whorled ontogenetic sequences, including those of subadult and mature specimens, are especially uncommon. Using methods published in 2021 to address the accessibility problem in sutural ontogenies, our paper represents the first comparisons of the fractal trajectories of ammonoids and includes several significant Jurassic ammonite families. We found that ammonite sutures did not steadily increase in complexity, as was previously suggested. Instead, juvenile ammonoids were more likely to exhibit ups and downs in their fractal ontogeny. We also found that pathologies must be considered as extremely common phenomena for ammonoids. Finally, we explored the ways that the unsteady ontogenetic trends of fractal complexity in ammonoids offer insights into the formation mechanism of ammonoid septa.
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