The morphology of fossil footprints is the basis of vertebrate footprint ichnology. However, the processes acting during and after trace fossil registration which are responsible for the final morphology have never been precisely defined, resulting in a dearth of nomenclature. Therefore, we discuss the concepts of ichnotaphonomy, ichnostratinomy, taphonomy, biostratinomy, registration and diagenesis and describe the processes acting on footprint morphology. In order to evaluate the morphological quality of tetrapod footprints, we introduce the concept of morphological preservation, which is related to the morphological quality of footprints (M-preservation, acronym MP), and distinguish it from physical preservation (P-preservation, acronym PP), which characterizes whether or not a track is eliminated by taphonomic and diagenetic processes. M-preservation includes all the morphological features produced during and after track registration prior to its study, and may be divided into substages (ichnostratinomic, registrational, taphonomic, stratinomic, diagenetic). Moreover, we propose an updated numerical preservation scale for M-preservation. It ranges from 0.0 (worst preservation) to 3.0 (best preservation); intermediate values may be used and specific features may be indicated by letters. In vertebrate footprint ichnotaxonomy, we regard the anatomy-consistent morphology and to a lesser extent the trackway pattern as the only acceptable ichnotaxobases. Only footprints showing a good morphological preservation (grade 2.0-3.0) are useful in ichnotaxonomy, whereas ichnotaxa based on poor morphological preservation (grade 0.0-1.5) are considered ichnotaphotaxa (nomina dubia) characterized by extramorphologies. We applied the preservation scale on examples from the Palaeozoic to the present time, including three ichnotaphotaxa and 18 anatomy-consistent ichnotaxa/morphotypes attributed to several vertebrate footprint producers. Results indicate the utility, feasibility and suitability of this method for the entire vertebrate footprint record in any lithofacies, strongly recommending its use in future ichnotaxonomic studies.

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This paper contains the first comprehensive ichnotaxonomic review of the Triassic tetrapod track record in Argentina, including previous accounts and new material recently discovered, and an analysis of its composition and stratigraphic distribution. Triassic footprints have been recorded from three basins: the Ischigualasto-Villa Unió n and Cuyo basins in north-west Argentina, and the Los Menucos depocentre in northern Patagonia. Most are in successions of Middle Triassic age; a lower number are from the Late Triassic, and there are two records from Early Triassic rocks. The known track types include: Brachychirotherium isp., cf. Brachychirotherium isp., Chirotherium barthii, Dicynodontipus ispp., Grallator isp., Rhynchosauroides isp., Rigalites ischigualastianus, Rigalites isp., Tetrasauropus isp., and bird-like, chirotheriid and unidentified tridactyl footprints. The ichnogenera Gallegosichnus Casamiquela, 1964, Calibarichnus Casamiquela, 1964, Palaciosichnus Casamiquela, 1964 and Stipanicichnus Casamiquela, 1975 are considered to be synonyms of Dicynodontipus (Hornstein, 1876). In addition, the abandonment of the following ichnogenera (and single ichnospecies) that are based on poorly preserved material is suggested: Ingenierichnus sierrai Casamiquela, 1964, Rogerbaletichnus aguilerai Casamiquela, 1964 and Shimmelia chirotheroides Casamiquela, 1964. At least eight Triassic ichnofaunas can be recognized. The most peculiar is that of the Late Triassic Los Menucos depocentre, which is characterized by the dominance of therapsid footprints (Dicynodontipus ispp.). The track assemblages from the Cuyo Basin display the highest ichnodiversity, with five footprint types.

Well-exposed Triassic rift strata from the Ischigualasto-Villa Unió n Basin (NW Argentina) include a 80 to ca 515 m thick lacustrine-dominated package that can be correlated across a half-graben using key stratigraphic surfaces (sequence boundaries, lacustrine flooding surfaces and forced regressive surfaces). The characteristics of the synrift lacustrine fill in different parts of the half-graben have been examined and the mechanisms controlling sedimentation inferred. A variety of sedimentary environments are recognized including; volcaniclastic floodplain, mildly saline lake and playa lake, offshore lacustrine, delta front to fluvial-dominated and wavedominated deltas, distributary and fluvial channel, and interdistributary bay. The succession can be divided into four stratigraphic sequences (SS1 to SS4), the oldest of which (SS1) contains volcaniclastic, fluvial and saline lake deposits; it is thickest close to the western border fault zone, reflecting more rapid subsidence here. Accommodation exceeded sediment and water input during SS1. The second and third sequences (SS2 and SS3) mark the onset of widespread lacustrine sedimentation, reflecting a balance between accommodation creation and water and sediment fluxes. Sequences SS2 and SS3 are represented by offshore meromictic lacustrine and deltaic deposits, the latter mostly sourced from the flexural and southern axial margins of the half-graben. The presence of stacked parasequences bound by lacustrine flooding surfaces is related to climatically induced lake-level fluctuations superimposed on variable rates of subsidence on the controlling rift border fault zone. The youngest sequence (SS4) is represented by the deposits of littoral lacustrine and shallow shelf deltas distinguished by a change in lithofacies, palaeocurrents and sandstone composition, suggesting a switch in sediment supply to the footwall margin to the NW. The change in the sediment source is related to reduced footwall uplift, the possible presence of a relay ramp and/or supply from a captured antecedent drainage network. During SS4, the rate of creation of accommodation was exceeded by the sediment and water discharge. The stratigraphic evolution of lacustrine strata in the half-graben was mainly controlled by tectonic processes, including subsidence rate and the growth and evolution of the border fault zone, but changing climate (inducing changes in water balance and lake level) and autocyclic processes (delta lobe switching) were also important.