The skeletal record of tyrannosaurids is well-documented, whereas their footprint record is surprisingly sparse. There are only a few isolated footprints attributed to tyrannosaurids and, hitherto, no reported trackways. We report the world’s first trackways attributable to tyrannosaurids, and describe a new ichnotaxon attributable to tyrannosaurids. These trackways are from the Upper Cretaceous (Campanian - Maastrichtian) of northeastern British Columbia, Canada. One trackway consists of three tridactyl footprints, and two adjacent trackways consist of two footprints each. All three trackways show animals bearing southeast within an 8.5 meter-wide corridor. Similarities in depth and preservation of the tyrannosaurid tracks indicate that these three trackways were made by track-makers walking concurrently in the same direction. These trackways add significantly to previous osteology-based hypotheses of locomotion and behavior in Tyrannosauridae by providing ichnologic support for gregariousness in tyrannosaurids, and the first record of the walking gait of tyrannosaurids.

A Late Pleistocene hominin tracksite has been identified in coastal aeolianite rocks on the Cape south coast of South Africa, an area of great significance for the emergence of modern humans. The tracks are in the form of natural casts and occur on the ceiling and side walls of a ten-metre long cave. Preservation of tracks is of variable quality. Up to forty hominin tracks are evident. Up to thirty-five hominin tracks occur on a single bedding plane, with potential for the exposure of further tracks. Five tracks are apparent on a second hominin track-bearing bedding plane. A number of individuals made the tracks while moving down a dune surface. A geological investigation at the site and stratigraphic comparison to published geochronological studies from this area suggest that the tracks are ~90 ka in age. If this is the case, the shoreline at the time would have been approximately 2 km distant. This is the first reported hominin tracksite from this time period. It adds to the relatively sparse global record of early hominin tracks, and represents the largest and best preserved archive of Late Pleistocene hominin tracks found to date. The tracks were probably made by Homo sapiens.

Tracks of small quadrupedal ornithischians with five manual and four pedal digits have been recorded from sedimentary rocks near the Late Jurassic-Early Cretaceous (TithonianBerriasian) boundary in NE Thailand and British Columbia. These are compared with larger tracks of gracile, quadrupedal ornithopods from the earliest Cretaceous of Spain and smaller tracks of a quadruped of unknown age from Zimbabwe. The Thai and Canadian tracks are similar to the Early Jurassic (Liassic) ichnogenus Anomoepus and the small ornithopod tracks from the Late Jurassic of Spain. They are the only known post-Liassic ornithischian tracks in which up to five discrete manus digit impressions are clearly visible. Based on strong heteropody (manus much smaller than pes) in all cases we infer an ornithopod trackmaker rather than another ornithischian. The scattered, but widespread earliest Cretaceous occurrence of this ichnotaxon, herein assigned to Neoanomoepus perigrinatus ichnogen. and ichnosp. nov., on the basis of type material from Canada, suggests that these hitherto unknown earliest Cretaceous ichnofaunas may represent a radiation of small basal ornithopods (pes length less than 15 cm), appearing before the widespread radiation of large ornithopods (pes length up to 60 cm or more) later in the Neocomian (Valanginian-Barremian), Aptian-Albian and Late Cretaceous. The primitive condition of the trackmaker is indicated by the pedal and manual morphology, which consists of four and five digits respectively that are not enclosed by well-developed fleshy padding or integument. In contrast, all larger Cretaceous ornithopod tracks, mostly from post-Berriasian strata, have only three pedal digits enclosed in fleshy pads and a manus in which all functional digits are reduced and enclosed by substantial flesh.

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