The pa Âramo is a high-altitude ecosystem of the northern Andes. The vegetation is continuous, with grasses as the dominant groundcover. Because of their high water retention, pa Âramos play a fundamental role in water availability for all the population of the inter-Andean valleys. There are many studies of this speci®c ecosystem, but very few are focussed on overgrazing and its effect on vegetation and soil properties.Intensive grazing started less than 20 years ago and was studied in a representative area in the western Cordillera of central Ecuador covered by recent volcanic ash deposits. Intensive sheep grazing has led to a strong decline in the number of plant species, the replacement of the tussock grass vegetation by a short carpet grass vegetation, and an increase of bare land. In that area, the upper 50 cm of Andisols are deeply affected by a convergent decrease of Al and Fe oxalate and pyrophosphate in soil extracts, carbon contents decrease from 100 g kg ±1 to less than 50 g kg ±1 in the humid zone, from 70 to 40 g kg ±1 in the dry zone and a reduction of water-retention capacity at ±33 kPa matrix potential from 800 g kg ±1 to 200 g kg ±1 in humid zones, from 350 to less than 100 g kg ±1 in drier areas. They showed also a decrease in the macrostructure and the development of a highly water repellent microstructure. All these important transformations favour the development of aeolian erosion in dry areas, runoff on bare surfaces, and gully erosion on slopes. The role of the pa Âramo in water-regulation of this ecosystem seems to have been adversely affected for the future.
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The impact of extensive livestock farming on the physical and chemical characteristics of the volcanic soils and on the nutrient status of green plant tissues of neotropical alpine grasslands (p~amo) is studied. Soil and plant samples were taken over a one-year period at five sites with different agricultural (grazing and burning) management. In the undisturbed pAramo ecosystem, soil moisture (50-250%) and organic matter content are high (7-27%) and decomposition (11-35% yr -1) and element concentrations are low. Low temperatures (max < 10°C) and phosphorus fixation by the soil (5 mg P g-1 soil) determine the low mineralization and turn-over rates.Multivariate analysis of laboratory results indicates that the season of sampling and the agricultural practice are the most important explanatory factors for variation of soil characteristics. After long-term heavy grazing, soils have a higher bulk density and a lower moisture content. The outcome of a litterbag experiment confirms the hypothesis of higher decomposition rates at grazed sites. In the intermediate (wet-dry) season, conditions were somewhat better for plant growth but the system remained nutrient limited. Surprisingly, no relation between soil density, moisture or carbon content and concentrations of available nutrients in the soil is found. This is supported by the rather uniform nutrient concentrations in green plant tissue among the sites. It is concluded therefore that the effect of burning and grazing on p~amo soils is principally restricted to physical characteristics, and that differences in chemical characteristics of the soil do not cause differences in vegetation structure between grazed, burned and undisturbed sites.
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