To ascertain whether browsing or hydrologic conditions influence the physiological performance of Salix and whether Salix and graminoids (Carex) use and possibly compete for similar water resources, we quantified the in situ seasonal patterns of plant water and carbon relations over three growing seasons. Our studies were designed to address the physiological factors which may be responsible for poor woody plant regeneration in montane riparian habitats of Rocky Mountain National Park, Colo. As these systems act to insure the delivery of fresh water to downstream users, the maintenance of their integrity is critical. We quantified plant water potentials, instantaneous rates of carbon fixation, leaf carbon isotope discrimination (Δ), leaf nitrogen content and water sources using stable isotopes of water (δO). The carbon and water relations of Salix were significantly affected by winter browsing by elk and in some cases by landscape position with regard to proximity to active streams. Winter browsing of Salix by elk significantly increased summer plant water potentials and integrative measures of gas exchange (Δ), though browsing did not consistently affect instantaneous rates of photosynthesis, leaf nitrogen or the sources of water used by Salix. No effect of experimental manipulations of surface water conditions on Salix physiology was observed, likely due to the mesic nature of our study period. Using a two-member linear mixing model, from δO values we calculated that Salix appears to rely on streams for approximately 80% of its water. In contrast, the graminoid Carex derives almost 50% of its water from rainfall, indicating divergent water source use by these two life forms. Based on these findings, winter browsing by elk improved Salix water balance possibly by altering the shoot to root ratio which in turn leads to higher water potentials and higher degrees of season-long gas exchange, while experimental damming had in general no effect on the physiological performance of Salix plants. In addition, as the water sources of Salix and Carex were significantly different, competition for water may not influence the growth, development, and regeneration of Salix. Thus, under the conditions of our study, herbivory had a positive effect on the physiological performance of Salix, but it is still unclear whether these changes in physiology transcend into improved Salix regeneration and survivorship. However, under drier environmental conditions such as lower snowpacks and lower stream flows, the browsing resistance of Salix and ecosystem regeneration may be greatly hindered because the reliance of Salix on stream water makes it vulnerable to changes in surface water and hydrological conditions.
An extensive system of NW striking faults constitutes a major tectonic feature of the Coastal Cordillera in northern Chile, but fundamental questions remain about timing and kinematics of these structures. We present new geologic mapping and geochronology that provide insight into the structural evolution and tectonic significance of the Taltal fault system (TFS). The TFS displaces the Early Cretaceous arc-parallel Atacama fault system (AFS) with~10.6 km cumulative offset across a~15 km wide zone. Brittle fault data demonstrate that the TFS is vertical to steeply NE dipping with an average sinistral slip vector plunging 11°from the NW, compatible with E-W shortening. Two late Early Cretaceous dikes cut the AFS but are cut by TFS faults, and synkinematic calcite on a TFS strand yielded a U-Pb calcite date of 114.1 ± 7.0 Ma. These data demonstrate that the AFS was abandoned and deformation (re) initiated on the TFS between~114-107 Ma, with continued slip after intrusion of the Tropezón (~110 Ma) and Librillo (106-101 Ma) plutonic complexes. Emplacement of a~146 Ma rhyolite dike along the main Taltal fault and 141 ± 11 Ma calcite mineralization in the fault core suggests that a precursor structure influenced magma emplacement and fluid flow in the Late Jurassic/Early Cretaceous, supporting the hypothesis that the TFS reactivated long-lived inherited crustal weaknesses. The Early Cretaceous shift from arc-parallel shear to slip on the TFS and E-W shortening shortly preceded migration of the magmatic arc and records a change in the Chilean margin subduction dynamics.
Displacement estimates along the Atacama fault system (AFS), a crustal-scale sinistral structure that accommodated oblique convergence in the Mesozoic Coastal Cordillera arc, vary widely due to a lack of piercing points. We mapped the distribution of plutons and mylonitic deformation along the northern ∼70 km of the El Salado segment and use U-Pb geochronology to establish the slip history of the AFS. Along the eastern branch, mylonitic fabrics associated with the synkinematic ∼134–132 Ma Cerro del Pingo Complex are separated by 34–38 km, and mylonites associated with a synkinematic ∼120–119 Ma tonalite are separated by 20.5–25 km. We interpret leucocratic intrusions to be separated across the western branch by ∼16–20 km, giving a total slip magnitude of ∼54 ± 6 km across the El Salado segment. Kinematic indicators consistently record sinistral shear and zircon (U-Th)/He data suggest dip-slip motion was insignificant. Displacement occurred between ∼133–110 Ma at a slip rate of ∼2.1–2.6 km/Myr. This slip rate is low compared to modern intra-arc strike-slip faults, suggesting (1) the majority of lateral slip was accommodated along the slab interface or distributed through the forearc or (2) plate convergence rates/obliquity were significantly lower than previously modeled.Supplementary material including full U-Pb, (U-Th)/He, petrographic, and structural data with locations is available at https://doi.org/10.6084/m9.figshare.c.5262177.
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