Military life is characterized by regular transitions; thus, parents are positioned to serve as stable protective factors for adolescents. We investigated a theory-informed model that assessed direct and indirect relationships between parental relationship quality, parental behaviors, and adolescent depressive symptomatology using cross-sectional data of military families in the United States (US). Participant families (N = 229), recruited via convenience sampling to take a computer-based survey, included an active duty father, his spouse, and an adolescent. Mother's couple relationship quality was indirectly linked to adolescent depressive symptoms through maternal warmth. Conversely, father's couple relationship quality was indirectly linked to adolescent depressive symptoms via paternal hostility. In other words, parental couple relationship quality was indirectly related to adolescent depressive symptoms, but this relationship differed by parent (i.e., warmth for mothers and hostility for fathers). Findings were similar for adolescent boys and girls.This study builds on tenets of family systems theory and the spillover hypothesis and incorporates suppositions regarding the gendered nature of parenting (Repetti, 1987) to examine the links between couple relationship quality, parental behaviors (particularly parental warmth and parental hostility), and adolescent mental health with a multi-informant study of military families. The relational health of the marital/couple subsystem is theoretically and empirically linked to the well-being of children within the family (e.g., Stroud et al., 2015), but less is known about what factors within the family system link couple relationship quality to the mental health of dependents. This study highlights parental behaviors as the linking mechanism between couple relationship quality and adolescent mental health. Evidence suggests that challenges in couple relationships (e.g., high levels of conflict) disrupt maternal and paternal parenting behaviors, such as involvement,