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The evolution of Mediterranean landscapes during the Holocene has been increasingly governed by the complex interactions of water and human land use. Different land-use practices change the amount of water flowing across the surface and infiltrating the soil, and change water's ability to move surface sediments. Conversely, water amplifies the impacts of human land use and extends the ecological footprint of human activities far beyond the borders of towns and fields. Advances in computational modelling offer new tools to study the complex feedbacks between land use, land cover, topography and surface water. The Mediterranean Landscape Dynamics project (MedLand) is building a modelling laboratory where experiments can be carried out on the long-term impacts of agropastoral land use, and whose results can be tested against the archaeological record. These computational experiments are providing new insights into the socio-ecological consequences of human decisions at varying temporal and spatial scales.

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The emergence of coupled natural and human landscapes marked a transformative interval in the human past that set our species on the road to the urbanized, industrial world in which we live, and enabled technologies and social institutions responsible for human-natural couplings in domains beyond rural, agricultural settings. The Mediterranean Landscape Dynamics Project (MedLand) is studying the interacting social and biophysical processes associated with these novel socioecological systems and their long-term consequences using a new form of 'experimental socioecology' made possible by recent advances in computation. We briefly describe the MedLand modeling laboratory, a hybrid simulation environment that couples models of smallholder farming and herding, landscape evolution, and vegetation change managed through an interaction model. We then review three examples of experimental socioecology carried out in this laboratory. These offer new insights for scale-dependent thresholds in agropastoral productivity, long-term sustainability of alternative land-use strategies, and identifying signatures of human and climate-driven landscape dynamics. We conclude with an overview of new directions for this interdisciplinary research on Anthropocene human-earth systems, including: modeling more diverse decision-making strategies for land-use, developing more sophisticated models of vegetation dynamics and fire ecology, and generating digital proxy data for more robust model validation against the empirical record.