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A characteristic feature of the Moldanubian part of the central European Variscides is late-orogenic high-T/low-P metamorphism. Its past history and the possible reasons for this metamorphism are highlighted by the tectonometamorphic development at the southeastern margin of the Bohemian massif. During the Variscan orogeny, at ca. 340 Ma, two different crustal segments were juxtaposed by thrusting (the Drosendorf unit on top of the Monotonous unit). This probably marks a collisional event that is widespread in the southeastern Moldanubian zone. The collision was followed by crustal uplift accompanied by strong heating in the lowermost structural unit (Monotonous unit). During the subsequent orogenic collapse, the Moldanubian nappe pile was thrust over parts of the Moravo-Silesian terrane. A late stage of crustal extension under greenschist-facies conditions is linked with pluton emplacement. In general, magmatic underplating as well as delamination of the lithospheric mantle explains the high-T/low-P metamorphism and the largescale plutonism in the southeastern Moldanubian zone.