We study theoretically the collective dynamics of particles driven by an optical vortex along a circular path. Phase equations of N particles are derived by taking into account both hydrodynamic and repulsive interactions between them. For N = 2, the particles attract with each other and synchronize, forming a doublet that moves faster than a singlet. For N = 3 and 5, we find periodic rearrangement of doublets and a singlet. For N = 4 and 6, the system exhibits either a periodic oscillating state or a stable synchronized state depending on the initial conditions. These results reproduce main features of previous experimental findings. We quantitatively discuss the mechanisms governing the non-trivial collective dynamics.

The thermal properties of cells were determined to depend on both temperatures and local heating frequencies using on-chip-integrated microthermistors.