Seedlings in regenerating layer are frequently attacked by herbivorous insects, while the combined effects of defoliation and shading are not fully understood. In the present study, two Leguminosae species (Robinia pseudoacacia and Amorpha fruticosa) were selected to study their responses to combined light and defoliation treatments. In a greenhouse experiment, light treatments (L+, 88% vs L−, 8% full sunlight) and defoliation treatments (CK, without defoliation vs DE, defoliation 50% of the upper crown) were applied at the same time. The seedlings’ physiological and growth traits were determined at 1, 10, 30, and 70 days after the combined treatment. Our results showed that the effects of defoliation on growth and carbon allocation under high light treatments in both species were mainly concentrated in the early stage (days 1–10). R. pseudoacacia can achieve growth recovery within 10 days after defoliation, while A. fruticosa needs 30 days. Seedlings increased SLA and total chlorophyll concentration to improve light capture efficiency under low light treatments in both species, at the expense of reduced leaf thickness and leaf lignin concentration. The negative effects of defoliation treatment on plant growth and non-structural carbohydrates (NSCs) concentration in low light treatment were significantly higher than that in high light treatment after recovery for 70 days in R. pseudoacacia, suggesting sufficient production of carbohydrate would be crucial for seedling growth after defoliation. Plant growth was more sensitive to defoliation and low light stress than photosynthesis, resulting in NSCs accumulating during the early period of treatment. These results illustrated that although seedlings could adjust their resource allocation strategy and carbon dynamics in response to combined defoliation and light treatments, individuals grown in low light conditions will be more suppressed by defoliation. Our results indicate that we should pay more attention to understory seedlings’ regeneration under the pressure of herbivorous insects.
Based on the study of cores, thin sections and outcrops, the sedimentary facies of the Lower Cambrian carbonate rocks in the northern Tarim Basin can be divided into four types: restricted platform, open platform, ramp and basin. Based on the lithologic analysis of thin sections, two-dimensional seismic data interpretation and an isopach map of the Lower Cambrian Xiaoerbulake Formation in the study area, seven sedimentary facies of carbonate rocks were identified, including inner platform depression, shoal, intershoal sea, platform margin, gypsum salt lake, ramp and basin. The depositional model of the Lower Cambrian Xiaoerbulake Formation in the northern Tarim Basin is constructed based on this integrated research. The topset, foreset and bottomset of oblique progradational reflections are interpreted as the platform margin beach, ramp and basin environments, respectively. The thicker area with micritic dolomite as the dominant lithology is interpreted as a platform depression. The low-amplitude hummocky reflections are interpreted as shoals that consist of a variety of granular dolomite and algal dolomite. The thinner uplifted area with gypsum and dolomite present in cores is interpreted as a restricted platform with a gypsiferous dolomite tidal flat or lake environment. Well LT1 which was drilled recently in the platform margin and ramp region, as indicated by seismic progradational clinoform reflections, has produced high yields of oil and gas. Supported by the above research results, the map of the lithofacies and paleogeography of the Xiaoerbulake Formation in the northern Tarim Basin was recompiled.
The Ordos Basin contains abundant coal resources in the Upper Paleozoic strata. They are shallowly buried or exposed in the periphery of the basin, forming recoverable coal mines and coalbed methane resources. For the deeply buried coal seams of the basin, however, due to complex changes and difficult mining, less attention was paid before. In recent years, with the exploitation of tight gas related to coal source rocks, a large amount of drilling and seismic data has been accumulated in the southeastern Ordos Basin, which makes it possible to study the distribution change and factors controlling Upper Paleozoic coal seams. Delineation of deep coal development and distribution is an important basis for coal, coalbed methane and related tight gas resources in the Upper Paleozoic. We use core, logging and seismic data to analyze the coal geophysics, thickness variation, paleotopography and sedimentary facies in the southeastern Ordos Basin. The lower part of the Shanxi Formation (Fm.) and the upper part of the Taiyuan Fm. in the southeastern part of the basin are characterized by low natural gamma ray, low density, high resistivity and high sonic logging values. As the coal velocity is very low and alternates with the high-velocity Taiyuan Fm. and Ordovician limestones, the two sets of coals below and above the Taiyuan limestones form an extremely strong-amplitude reflection in the seismic profile. The two sets of coals are cumulatively 1–10 m thick, averaging 2.4 m, and 4 m is the maximum induvial thickness. The individual coal seams thicker than 2 m cover an area of approximately 2500 km2, and the burial depths of the coal seam range from 2500 to 3000 m. The thick coal seams of the Taiyuan Fm. are mainly distributed proximally in the Ordovician paleokarst depression, while the Ordovician paleokarst depression without incised origin was deposited by distal coeval limestone rocks. There is a complementary relationship between the coal seams and the thickened zone of distributary channel sandstone in the Shanxi Fm.
Coal-measure source rocks are generally developed in marsh facies under a humid climate and are rarely reported in a carbonate platform or a mixed platform. Carboniferous seawater intruded from west to east in the Tarim Basin, and mixed platform deposits of interbedded mudstone and carbonate developed in the southwest of the basin. In recent years, with the deepening of the exploration, nearly 20 m coal seams and carbonaceous mudstone source rocks have been found in the Carboniferous lagoon’s tidal-flat background. The hydrocarbon generation potential, development, and distribution of these coal-measure source rocks have become an important issue for oil and gas exploration. Coal seams and carbonaceous mudstones were found in the Carboniferous formation of wells BT5, BT10, and Lx2. The hydrocarbon prospect, development, and distribution characteristics of these coal formations have become an important research topic. The authors conducted organic geochemical tests and analyses of core and samples drill cuttings from multiple wells in the study area, combined with research focused on the identification and distribution of coal seams, dark mudstones, and depositional facies via logging cross plots of different lithology and 3D seismic inversion. The results show that coal-measure source rocks in the BT5 well are related to the set of delta-lagoon sedimentary systems widely developed in the Carboniferous Karashayi Formation. The maximum cumulative thickness of coal-measure source rocks is about 20 m, with total organic carbon (TOC) contents of 0.15–60%, kerogen types II2-III, and vitrinite reflectance (Ro) values of 0.78–1.65%. The rocks have generally low maturity in the northwestern area and high maturity in the southeastern area, and the maturity changes as the burial depth changes. The effective hydrocarbon source rocks such as coal, carbonaceous mudstone, and dark mudstone all show acoustic time (AC) greater than 300 µs/m, and density (DEN) less than 2.3 g/cm3, but possess different gamma ray (GR) values. The GR value is less than 75 API for coal, between 75–100 API for carbonaceous mudstone, and greater than 100 API for dark mudstones. The distribution of source rocks can be identified in the area between the wells according to a 3D seismic inversion impedance (IMP) of less than 7333 m/s·g/cm3. The development and controlled factors of coal-measure source rocks of delta facies in the mixed platform have a significant role for oil and gas exploration of Upper Paleozoic in this area. The coal measure and sandstones of delta in the Carboniferous are expected to form self-generation and self-storage pools in this area.
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