Abstract-Several continuous-time tree-grass competition models have been developed to study conditions of long-lasting coexistence of trees and grass in savanna ecosystems according to environmental parameters such as climate or fire regime. In those models, fire intensity is a fixed parameter while the relationship between woody plant size and fire-sensitivity is not systematically considered. In this paper, we propose a mathematical model for the tree-grass interaction that takes into account both fire intensity and size-dependent sensitivity. The fire intensity is modeled by an increasing function of grass biomass and fire return time is a function of climate. We carry out a qualitative analysis that highlights ecological thresholds that summarize the dynamics of the system. Finally, we develop a non-standard numerical scheme and present some simulations to illustrate our analytical results.
The savanna biome encompasses a variety of vegetation physiognomies that traduce complex dynamical responses of plants to the rainfall gradients leading from tropical forests to hot deserts. Such responses are shaped by interactions between woody and grassy plants that can be either direct, disturbance-mediated or both. There has been increasing evidence that several vegetation physiognomies, sometimes highly contrasted, may durably coexist under similar rainfall conditions suggesting multi-stability or at least not abrupt transitions. These fascinating questions have triggered burgeoning modelling efforts which have, however, not yet delivered an integrated picture liable to furnish sensible predictions of potential vegetation at broad scales. In this paper, we will recall the key ecological processes and resulting vegetation dynamics that models should take into account. We will also present the main modelling options present in the literature and advocate the use of minimalistic models, capturing only the essential processes while retaining sufficient mathematical tractability and restricting themselves to a minimal set of parameters assessable from the overall literature.
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