Strawberry (Fragaria ×ananassa Duch.) fruit have a unique, highly desirable flavor and are one of the most popular summer fruits. Sugars, acids, and aroma volatiles contribute to the characteristic strawberry flavor, which is dependent on the proper balance of these chemical constituents. While sugars and acids are responsible for the sweetness and tartness of the fruit, aroma volatiles provide the unique, fruity flavors that characterize a fresh strawberry. The aroma of fresh strawberries is dependent on many factors. The large genetic variability in the nature of strawberry aroma results in differences in flavor among cultivars. In addition, the aroma changes dramatically during fruit ripening after harvest; therefore, it is important to preserve and enhance the ripe fruit aroma during postharvest handling. The loss of this desirable aroma or the development of objectionable aromas reduces the quality and marketability of fresh strawberries. In this review we will discuss the chemical nature of strawberry aroma and how it is affected by various factors, including cultivar, maturity, and postharvest environment. CHEMICAL COMPOSITION OF AROMA Volatile chemicals are responsible for the aroma and contribute to the flavor of fresh strawberries. These compounds comprise only 0.01% to 0.001% of the fruit fresh weight but have a major effect on its quality (Buttery, 1981). Fresh strawberries produce numerous volatile compounds; as many as 360 have been isolated (Latrasse, 1991) including esters, aldehydes, ketones, alcohols, terpenes, furanones, and sulfur compounds (McFadden et al., 1965). Esters are quantitatively and qualitatively the most abundant class of these compounds; 131 different ones have been identified in strawberry aroma (Latrasse, 1991). Esters provide the fruity and floral notes and they comprise from 25% to 90% of the total volatiles in fresh ripe fruit (Douillard and Guichard, 1990; Ito et al., 1990; Pyysalo et al., 1979; Schreier, 1980). Other classes of compounds, which may comprise up to 50% of strawberry volatiles, include aldehydes (Schreier, 1980) and furanones (Larsen and Poll, 1992). Alcohols account for as much as 35% of the volatiles, but normally contribute little to strawberry aroma (Larsen and Watkins, 1995b). While terpenes normally comprise <10% of strawberry volatiles and sulfur compounds <2%,
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