There is widespread concern that fire exclusion has led to an unprecedented threat of uncharacteristically severe fires in ponderosa pine (Pinus ponderosa Dougl. ex. Laws) and mixed-conifer forests of western North America. These extensive montane forests are considered to be adapted to a low/moderate-severity fire regime that maintained stands of relatively old trees. However, there is increasing recognition from landscape-scale assessments that, prior to any significant effects of fire exclusion, fires and forest structure were more variable in these forests. Biota in these forests are also dependent on the resources made available by higher-severity fire. A better understanding of historical fire regimes in the ponderosa pine and mixed-conifer forests of western North America is therefore needed to define reference conditions and help maintain characteristic ecological diversity of these systems. We compiled landscape-scale evidence of historical fire severity patterns in the ponderosa pine and mixed-conifer forests from published literature sources and stand ages available from the Forest Inventory and Analysis program in the USA. The consensus from this evidence is that the traditional reference conditions of low-severity fire regimes are inaccurate for most forests of western North America. Instead, most forests appear to have been characterized by mixed-severity fire that included ecologically significant amounts of weather-driven, high-severity fire. Diverse forests in different stages of succession, with a high proportion in relatively young stages, occurred prior to fire exclusion. Over the past century, successional diversity created by fire decreased. Our findings suggest that ecological management goals that incorporate successional diversity created by fire may support characteristic biodiversity, whereas current attempts to “restore” forests to open, low-severity fire conditions may not align with historical reference conditions in most ponderosa pine and mixed-conifer forests of western North America.

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Subalpine spruce (Picea) – fir (Abies) forests occur throughout the Cordillera of western North America. A variety of alternative silvicultural systems to clear-cutting are being investigated in these high-elevation forests in terms of their impacts on ecosystem components. We tested the hypotheses that abundance, reproduction, and survival of populations of (i) southern red-backed voles (Clethrionomys gapperi Vigors) will decline, (ii) long-tailed voles (Microtus longicaudus Merriam) and northwestern chipmunks (Tamias amoenus J.A. Allen) will increase, and (iii) deer mice (Peromyscus maniculatus Wagner) will be similar, with respect to decreasing levels of tree retention. Small mammal populations were live-trapped from 1994 to 1998 in replicated sites of uncut forest, single tree selection, 0.1-ha patch cuts, 1.0-ha patch cuts, a 10.0-ha clearcut, and edges in an Engelmann spruce (Picea engelmannii Parry ex Engelm.) – subalpine fir (Abies lasiocarpa (Hook.) Nutt.) forest near Sicamous, British Columbia, Canada. Clethrion omys gapperi dominated the small mammal community, reaching a peak abundance of 80 animals/ha with mean values ranging from 31 to 50 animals/ha. Over the 4 postharvest years, abundance, reproduction, and survival of C. gapperi populations were consistently similar among uncut forest, single tree selection, and patch cut treatments compared with clear-cut sites where this species declined to extirpation. Microtus longicaudus and T. amoenus occurred predominantly in clearcut sites, whereas P. maniculatus was present in low numbers on all sites. With respect to small mammals, partial harvesting systems appear to provide a means for combining timber extraction with maintenance of mature forest habitat in these subalpine ecosystems.

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