Structures previously suggested to be footprints or tracks, but which have been little described, were observed within Quaternary inland dunes of the European Sand Belt. Excavation of these in several localities of the eastern part of the belt in Poland reveal that they are hoofprints of various ungulate mammals, both wild and domestic, as well as footprints of humans, ranging from potentially as early as the fifth to sixth century to the 19th century or later. Trample grounds and horizons with high densities of tracks are associated with other signs of intensifying usage of dune habitats for pasture and farming, including constrained movement of livestock, relatively well-developed paleosols, and evidence of ploughing. The presence of such abundant life traces not only reflects the presence of individual animals and people in an area, but sheds light on the potential processes and feedbacks that the tracemakers were involved in, in terms of maintaining and modifying the habitats themselves, which were dynamic and changeable. Footprints and their associated sedimentological records can be integrated with archaeological and historical data to analyse anthropogenic influence through time on physical landscapes and on the wild biota.

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The earwigs, Dermaptera, are a group of insects which have been present since the Mesozoic. They have a relatively sparse fossil record, yet their life activities on and in soil or sediment leave traces with the potential for long-term preservation. These may include some burrows seen in Quaternary dunes and other sandy substrates. The well-known, cosmopolitan, sand-dwelling species Labidura riparia is examined as a potential model and reference for dermapteran tracemakers there and elsewhere in the geological record, through experimentally produced shelter burrows and trackways from wild-caught, laboratory-raised specimens. Shelter burrows were typically U-shaped with a pair of surface entrances, and these U-shapes could be additionally modified into Y-shapes or linked together to form a network. Trackways of L. riparia generally resembled those of other insects but may show features consistent with dermapteran anatomy such as tail-drag impressions produced by cerci.