Global terrestrial ecosystems take up around one-third of human emission of CO 2 (Friedlingstein et al., 2020). With recent climate change, the seasonality of terrestrial ecosystems, a sensitive indicator of biosphere-climate interactions, has been reported to be changing (Piao et al., 2007;Seddon et al., 2016). Longer growing seasons have been associated with increased carbon uptake by many ecosystems (Keenan et al., 2014), but the dominant controls of the growing season length remain relatively poorly understood.The start of growing season (SOS) has been found to have advanced in most ecosystems and is known to be largely controlled by air temperature in temperate and boreal ecosystems (Chmielewski

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