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The well-preserved Tvären crater is noteworthy for being one of a small number of Early and Middle Ordovician impact structures formed in a marine environment. It is demonstrated to be an impact structure by the presence of a breccia lens, consisting of crystalline basement rocks, and shocked quartz. The breccia lens formed under dry-hot conditions after expulsion of sea-water by the impact. Resurging sea-water thereupon deposited a positively graded, 60 m thick turbidite-like unit. This graded resurge deposit is a previously unknown feature, to be expected in open-sea impacts. Breccia in the lower part of this graded deposit contains fragments of a remarkably complete orthoceratite limestone succession that existed at the site of impact, resting on non-lithified sand of probably Early to earliest Middle Cambrian age. A sedimentary succession was deposited inside the crater at depths decreasing from more than 200 m in the initial stages to some 100 m at the time of deposition of the youngest preserved beds. The environment within the crater thus favoured deposition of an essentially complete stratigraphic succession with depth-controlled palaeoecology for a significant time interval after the impact. Whereas planktonic members, like graptolites and chitinozoa, are present throughout the post-impact succession, and asaphids, almost as persistent, became established at an early date, burrowers were somewhat reluctant to enter and remopleuridids and small strophomenids came in at a late stage. We suggest as a result of this study that structures formed by impact may offer unique information about the palaeogeology and palaeoenvironment of the region hit by the impact.

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-The chitinozoan biostratigraphy of four Ordovician impact craters has been investigated. Three of these (Tviiren, Kardla and Lockne) contain complete sequences of early Caradoc agc, while the Granby crater contains rocks of late Arenig age yielding two bentonitic horizons at their top. Chitinozoans, together with graptolites and other planktic organisms, were the first to invade the craters after the impact event. It has therefore been possible to date the impact events with a precision of less than one million ycars. An immigration of graptolites from Australia during the late Arenig corresponds to an immigration of chitinozoans from Gondwana at this time. Two stratigraphically important taxa, not previously described or discussed, Lagenochitina sp. A aff.

The taxonomy and distribution of Chitinozoa from Silurian (Llandovery) and Lower Devonian strata are reported from the Paraná Basin in southern Brazil and eastern Paraguay. The preCarboniferous sequences of the Paraná Basin in this area are present in three sub-basins viz., the Alto Garças (north) and Apucarana (south) sub-basins in Brazil, and the "East Paraguay Sub-basin" in east Paraguay. There is more similarity in the lithology between the Alto Garças and "East Paraguay" subbasins, than between the former and the Apucarana sub-basin. Llandoverian and Lochkovian to Pragian beds are present in all sub-basins. So far, no Emsian beds have been found in outcrops from the north-northwest margin of the Alto Garças Sub-basin, and no early Emsian beds in the outcrops on the northeast margin. Furthermore, Emsian beds could not be identified from the "East Paraguay Sub-basin" in the present study. The Early Devonian sequence is more complete in the Apucarana Sub-basin. It seems that the Apucarana Sub-basin endured a different evolution compared to the two other sub-basins during the Ordovician, Silurian and lower Devonian. Of the 39 chitinozoan species encountered, 24 species are left in open nomenclature, and the following five species are newly described: Ancyrochitina paranaensis, Angochitina daemoni, Sphaerochitina silurica, Spinachitina harringtoni, and Spinachitina wolfarti. A chitinozoan biozonation, with five zones and three subzones, is proposed for the investigated interval, and compared with the spore zonation. In addition we infer that the Furnas Formation could correspond to two chitinozoan zones found in Bolivia and Argentina (zones of Urochitina loboi and Angochitina aff. A. comosa), ranging in age from late Lochkovian to earliest Pragian. The zones are, from oldest to youngest: total range zone of Belonechitina postrobusta (upper Rhuddanian); total range zone of Conochitina elongata (Aeronian); concurrent range subzone of Spinachitina wolfarti n. sp. and Plectochitina sp. A (lower Aeronian); concurrent range subzone of Conochitina proboscifera and Spinachitina harringtoni n. sp. (upper Aeronian s.l.); concurrent range subzone of Conochitina proboscifera and Desmochitina cf. D. densa (lower Telychian s.l.); concurrent range zone of Salopochitina monterrosae and Conochitina proboscifera (upper Telychianlower Sheinwoodian); total range zone of Urochitina loboi (upper Lochkovian); total range zone of Angochitina aff. A. comosa (lowermost Pragian); total range zone of Ramochitina magnifica (Pragian s.l.), and total range zone of Ancyrochitina parisi (upper Emsian).