Plant pathogens pose a serious and continuous threat to global agricultural production and food security (Varden et al., 2017). Pathogens, including bacteria, fungi, oomycetes, viruses, nematodes, as well as feeding insects, subvert key host processes by suppressing host innate immunity and manipulating host nutrient supply, leading to disease. In most cases, pathogens achieve this goal by delivering virulence effector proteins into host cells, where the effectors attack the host innate immune system through various

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