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The oriental beech forests along the southern Caspian Sea shores of Iran are characterized by high biodiversity and high ecological value. However, there is little information concerning factors governing community composition and the presence and abundance of invasive plants. In the present study we aimed to evaluate the main drivers of community composition, diversity, and establishment of invasive plants in disturbed and undisturbed Fagus orientalis communities.We sampled 104 vegetation relevés in undisturbed and disturbed sites of the "Kheiroud Investigation Forest" in northern Iran and collected data on a large number of environmental variables concerning soil properties, light conditions, and topography. Data analysis consisted of a forward selection of signifi cant explanatory variables followed by partial CCA analyses. Additionally, we carried out indicator species analysis of groups of relevés affected by different disturbance types and analyzed effects of disturbance on species richness, diversity, and evenness using general linear models.CCA results showed that disturbance, soil properties, light conditions and elevation explained 22.6% of the total variation in fl oristic composition. Gap and gap + grazing were the most important disturbance types affecting community composition. Disturbance had signifi cant but rather weak effects on species richness, evenness, and the Shannon index. The most obvious effect of disturbance was on species composition, as revealed by signifi cant groups of indicator species for different disturbance types. Most of the indicator species in grazed sites were unpalatable, poisonous, and creeping species, which are avoided by cattle. Grazing led to decreased abundance of indicator species and species characteristic of old-growth Fagus forests. The most intensive disturbance type "harvesting + gap + grazing" was associated with invasion by native vascular plant species not typically found in beech forests (apophytes).

The development of societies, including spiritual development, is closely connected to forests. The larger interrelations among changing societies, transforming forest landscapes, and evolving spiritual values related to forests have yet to be extensively considered. Addressing this research gap is important to avoid the neglect of spiritual values in forest policy and management. Our exploratory study investigates spiritual values of forests from European and Asian perspectives, assessing 13 countries. Based on expert knowledge from 18 interdisciplinary experts, we first define forest spiritual values (forest spirituality). We then elaborate on the idea that forest spirituality evolves as societies and landscapes change, and propose a transition hypothesis for forest spirituality. We identify indicators and drivers and portray four stages of such a transition using country-specific examples. We find that during a first stage ("nature is powerful"), forest spirituality is omnipresent through the abundance of sacred natural sites and practices of people who often directly depend on forests for their livelihoods. An alternative form of spirituality is observed in the second stage ("taming of nature"). Connected to increasing transformation of forest landscapes and intensifying land-use practices, "modern" religions guide human-nature interrelations. In a third stage ("rational management of nature"), forest spirituality is overshadowed by planned rational forest management transforming forests into commodities for the economy, often focusing on provisioning ecosystem services. During a fourth stage ("reconnecting with nature"), a revival of forest spirituality (re-spiritualization) can be observed due to factors such as urbanization and individualizing spirituality. Our core contribution is in showing the connections among changing forest perceptions, changing land-use governance and practices, and changing forest spirituality. Increasing the understanding of this relationship holds promise for supporting forest policy-making and management in addressing trade-offs between spiritual values and other aspects of forests.

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