The Dolní Vě stonice-Pavlov-Milovice area (Czech Republic) on the slopes of the Pavlov Hills provides an opportunity for correlating the geomorphology of the Dyje River valley with Gravettian settlement patterns. Although the sites vary in size and complexity, they create a regular chain of strategic locations at elevations between 200 m and 240 m asl. In 2009, a road collapsed into deserted cellars inside the village of Milovice and revealed a complex of archaeological layers deep within loess, at an elevation of only 175 m asl. This paper presents an analysis of this atypical archaeological site location and compares the results with the other sites. We argue that this location allowed direct contact with mammoth herds concentrated on the floodplain, while the aquatic environment offered possibilities for gathering plants and fishing. This site represents a new aspect of organized settlement, hunting strategies, and short-distance human movements during the Gravettian within this area.